Chenopodium harae Sukhor., Phytotaxa 226(3): 290 (2015)

Sukhorukov, Alexander P., Liu, Pei-Liang & Kushunina, Maria, 2019, Taxonomic revision of Chenopodiaceae in Himalaya and Tibet, PhytoKeys 116, pp. 1-141 : 21-22

4. Chenopodium harae Sukhor., Phytotaxa 226(3): 290 (2015)

= Chenopodium pallidum sensu Sukhorukov (in Sukhorukov and Kushunina 2014), non Moquin-Tandon (1840).

Holotype.

NEPAL, [Dhaulagiri Zone], Mustang Prov. [Distr.], Annapurna Conservation area, Trekking route Jomosom-Muktinath, valley of Khali Gandaki river, between Jomosom and Kagbeni villages, riverside, 2700 m alt., 22 Sep 2009, A. Sukhorukov 566 (MW!).

Taxonomic notes.

In the latest revision of the Chenopodiaceae in Nepal ( Sukhorukov and Kushunina 2014), this species is cited as Chenopodium pallidum Moq., but a closer look at the authentic specimens in P has proved that all plants (collected in early blooming stage) are indeed Atriplex pallida , now synonymised with A. schugnanica Iljin ( Sukhorukov and Kushunina 2015).

Description.

Annual up to 40 cm with erect stem; lateral branches ascending if present. Leaves short-petiolate (petioles to 1 cm) with blades up to 2 × 1 cm, fleshy, ovate, slightly cuneate, green above, greenish below, dentate, margins often red. Inflorescence bracteate or aphyllous. Perianth green, enclosing the fruit; segments abaxially carinate. Fruit 1.3-1.5 mm, pericarp easily separating from the seed; papillae up to 25 µm. Seed blackish or reddish-black, surface striate with obtuse or acutish margin (immature seeds with acute margin) with small depression apically.

Habitat.

Disturbed areas; 1500-3200 m a.s.l.

Phenology.

Flowering: March-September; fruiting: April-November.

Distribution.

See Fig. 7 .

Specimens examined.

INDIA: Himachal Pradesh: Kinnaur Distr., Shongtong, 11 Jun 1972, K.P. Janandhanan 48358 (BSD); Simla Distr., Seoni, 2 Jun 1986, P.C. Pani 80903 (BSD);

NEPAL: Midwestern: Rapti Zone: [Rolpa Distr.] Phalabang, 4500 ft a.s.l., 24 Mar 1952, O. Polunin, W.R. Sykes & L.H.J. Williams 654 (BM, LE); Western: Dhaulagiri Zone: Mustang [Mustang Distr.], Jomosom to Kagbeni, 22 Sep 2009, A. Sukhorukov 566 (E00757737, MW); Myagdi Prov. [Distr.], nr Dana vill., 1500 m a.s.l., 11 May 2010, A. Sukhorukov 510 (MW).

General distribution.

Central and Northern Himalaya (India, Nepal, Pakistan [Gilgit: E!]).

Figures

Figure 7. Distribution map of Chenopodium harae (stars) and C. karoi (circles).