Ochterus marginatus marginatus ( Latreille, 1804 )

Kment, Petr, Carapezza, Attilio & Jindra, Zdeněk, 2020, Taxonomic catalogue of the family Ochteridae with description of Ochterus papaceki sp. nov. from Socotra Island and Tanzania (Hemiptera: Heteroptera), Acta Entomologica Musei Nationalis Pragae 60 (1), pp. 23-64 : 39-42

Ochterus marginatus marginatus ( Latreille, 1804) ( Fig. 7 )

Acanthia marginata Latreille, 1804: 242 –243 (original description).

SYNTYPE (S): France: Bordeaux (depository unknown, cf. POLHEMUS

1995a: 25).

Ochterus marginatus : LATREILLE (1807): 143 (new combination).

Pelogonus marginatus Latreille, 1809: 384 (new combination).

Pelogonus marginatus: LE PELETIER & SERVILLE (1825): 33 (redescription); DUFOUR (1833): 192–196, Pl. V: fig. 58 (diagnosis, distribution, illustration); BURMEISTER (1835): 203–204 (diagnosis, distribution); BRULLÉ (1836): 277–278 (redescription, distribution); BLANCHARD (1840): 93–94 (redescription, catalogue); AMYOT & SERVILLE (1843): 409 (diagnosis, distribution); HERRICH- SCHÄFFER (1849): 18, 23–24, Pl.CCXC: fig. 892 (diagnosis, catalogue, illustration); FIEBER (1861): 103 (catalogue); COSTA (1862): 331 (redescription); REUTER (1888): 721–722 (catalogue); OSHANIN (1909): 955–956 (catalogue); SINGH- -PRUTHI (1925): 184, Pl. XVIII:fig. 130 (male genitalia, illustrations).

Pelogonus marginatus (partim): STÅL (1866): 170–171 (redescription, catalogue); STÅL (1876): 137 (catalogue); DISTANT (1906): 14 (redescription).

Pelegonus marginatus: PUTON (1869): 38 (list); PUTON (1880): 209 (redescription, distribution).

Ochterus marginatus : HOFFMANN (1933): 249 –250 (catalogue); JACZEWSKI (1935): 482 (key to species); WU (1935): 557 (catalogue); KORMILEV (1971): 433 –436 (redescription, illustrations, variation); KORMILEV (1973): 2 –4, 8 (diagnosis, variability, illustrations); RIEGER (1976): 109 –188 (head and prothorax morphology, illustrations); RIEGER (1977): 213 –214 (differential diagnosis, illustration); TAMANINI (1979): 18, 45, 51 (key, redescription, illustrations); NIESER (1982): 11, 46 (illustration, distribution); BOULARD & COFFIN (1991): 58 –67 (adult and larval morphology, biology); NIESER & CHEN (1992): 9 (illustration of paramere); NIESER et al. (1994): 4, 20–21, 44 (key, illustrations); CHEN (1999): 9 –10 (diagnosis, illustrations); NIESER & CHEN (1999): 90 –91 (key, illustration); CHEN et al. (2005): 78 –79, 81, 413 (illustrations, bionomics, checklist); KANYUKOVA (2006): 46–47 (redescription, illustrations); POLHEMUS & POLHEMUS (2008): 286 (key to species); LINNAVUORI et al. (2011): 76 (photo); POLHEMUS & POLHEMUS (2012): 345 (key to species); POLHEMUS & POLHEMUS (2015): 207, 209–210 (key to species, illustrations); POPOV & HEISS (2014a): 187 –188 (illustrations).

Ochterus marginatus marginatus : JACZEWSKI (1934): 601 –605 (redescription, variability, illustrations); JACZEWSKI (1935):483 (key to species); POISSON (1957):24 –25 (redescription, illustrations); HOFFMANN (1941): 67 –69 (catalogue); POLHEMUS (1995a): 25 (catalogue); POLHEMUS & POLHEMUS (2012): 345 –347 (diagnosis, variability, illustrations); AUKEMA et al. (2013): 8 (catalogue); XIE & LIU (2013): 6 –7 (checklist); THIRUMALAI (2017): 15–16, 69 (catalogue); HAYASHI & MIYAMOTO (2018): 259, 261 (identification, illustration).

Pelogonus indicus Guérin-Méneville, 1843: 113 (original description, differential diagnosis). SYNTYPE (s): India: ‘le plateau des Neelgherries, dans le Coromandel’ [= Nilgiri Hills] (MZFN, see POLHEMUS 1995a: 25). Junior subjective synonym by MONTANDON (1910: 1).

Pelogonus indicus: STÅL (1876): 137 (catalogue).

Pelogonus armeniacus Kolenati, 1857: 454– 455. SYNTYPES: 6 (♀), Azerbaijan: Karabagh: ‘in littore lacuum alpestrium [= at coast of alpine lakes] Balloch-ghöll, Sullü-ghöll, Schamlogh-ghöll, Kaepesghöll in provincia Karabagh’ (ZMAS, probably additional syntypes in NHMW, see POLHEMUS 1995a: 25). Junior subjective synonym by KIRITSHENKO (1918: 172).

Pelogonus armeniacus: STÅL (1876): 137 (catalogue).

Pelogonus flavomarginatus Scott, 1874: 446 –447 (original description). HOLOTYPE:?sex, Japan (BMNH). Junior subjective synonym by JACZEWSKI (1934: 598, 605).

Pelogonus flavo-marginatus (incorrect subsequent spelling): STÅL (1876): 137 (catalogue).

Pelogonus flavomarginatus: MATSUMURA (1915): 103 (catalogue).

Ochterus marginatus flavomarginatus: MIYAMOTO (1961): 230 –231, 257, pl. 42: figs 235–237 (morphology of alimentary organs, illustrations); MIYAMOTO & LEE (1966): 398 (catalogue, distribution); TOMOKUNI (1981): 111 (distribution).

Ochterus flavomarginatus: HAYASHI (1995): 20 (distribution, habitat).

Salda rutherfordi Distant, 1915: 506 (original description). HOLOTYPE:?sex, Sri Lanka: ‘Peradeniya’ (BMNH). Junior subjective synonym; suspected synonymy suggested by W. E. China in JACZEWSKI (1934: 598) (see also POLHEMUS 1979: 112).

Pelogonus formosanus Matsumura, 1915: 103 –104 (differential diagnosis). SYNTYPES: 2 1 ♀, Taiwan: ‘Kagi’ [= Chiayi] and ‘Koshun’ [= Hengchun] (EIHU). Junior subjective synonymy by ESAKI (1926: 184); upgraded to subspecies by MIYAMOTO (1960: 78), synonymy confirmed by KORMILEV (1971: 433).

Ochterus formosanus: TAKAHASHI (1921): 119 –125 (biology); TAKAHASHI (1923): 67 –68 (biology).

Ochterus thienemanni (misidentification): NIESER & CHEN (1992): 12 (corrected by NIESER & CHEN 1999).

Material examined. Palaearctic Region: Europe: BULGARIA: K’rdzhali env., Arda river, 23.viii.1948, 4 1 ♀, M. Josifov det. ( NMPC); Momchilgrad, Djebel, 24.vi.1961, 6 3 ♀♀, L. Hobrlandt & Slouková lgt., P. Kment det. ( NMPC). FRANCE: Dép. du Var, [18]99, 1 ♀, no collector, L. Hoberlandt det. ( NMPC); Dept. Var, Le Lavandou, 1930, 1, J. Obenberger lgt., P. Kment det. ( NMPC); Mandelieu, Esleret [?], viii., 1 ♀, Bordeau lgt., J. Roubal det., P. Kment revid.( MMBC). GREE- CE: EAST MACEDONIA AND THRACE: Drama, Paranestion, 15.v.1937, 1, Bartoň lgt., P. Kment det. ( NMPC). ITALY: SICILY: Balestrate (PA), 7.vii.1972, 1 larva, 22.xi.1972, 1 larva,A. Carapezza lgt. & det. ( ACPI); Madonie, Petralia Sottana, 1000 m a.s.l., 9.vi.1972, 1 ♀, A. Carapezza lgt. & det. ( ACPI); Mezzoiuso (PA), 22.v.1977, 1, A. Carapezza lgt. & det. ( ACPI). Palermo Prov., Scillato env., Torrente Fichera, ca. 180 m a.s.l., 28.iv.2014, 1, J. Hájek & K. Kalíková lgt., P. Kment det.( NMPC). North Africa: ALGERIA: Ghardimaou, 19.–26.iv.[19]27, 2 ♀♀, Mařan lgt., P. Kment det. ( NMPC). Asia: AFGHANISTAN: LAGHMAN: Nengrahar prov., Laghman (30), 860 m a.s.l., 20.iii.1966, 1, Povolný & Tenora lgt., P. Kment det.( MMBC). Nuristan: Bashgultal [= Landai Sin valley], 1100 m a.s.l., 14.vii.1952, J. Klapperich lgt., P. Kment det. ( NMPC). CHINA: ANHUI: Dabieshan Mts., 65 km SW of Huoshan, 1400 m a.s.l., 23.vi.1998, 1 1 ♀, L. Bocák lgt., P.Kment det.( ZJPC). BEIJING: Pearl Lake [= Zhenzhu Lake], 70 km W of Beijing, 16.–18.vi.2000, 1, Z. Jindra lgt., P. Kment det. ( ZJPC). FUJIAN: Kuatun, 9.vii.1946, 1 ♀, Tschung Sen lgt., P. Kment det.( NMPC). SHAANXI: Qing Ling Shan Mts., road Baoji‒Taibai village, ca. 35 km S of Baoji, 18.vii.1998, 1 ♀, Z. Jindra lgt., P. Kment det. ( ZJPC). YUNNAN: Cheli env., 700 m a.s.l., 30.iv.1957, 1 L5, D. Panfilov lgt. ( NMPC); Lunan env., Stone Forest, 29.vii.1995, 1, Z. Jindra lgt., P. Kment det. ( ZJPC); 50 km SW of Mojiang, 1100 m a.s.l., 2.iv.1955, Kryzhanovskiy lgt., P. Kment det. ( NMPC); Xiaomengyang, 810 m a.s.l., 26.iii.1957, 2 1 ♀, S.-Y.Wang lgt., P. Kment det. ( NMPC); Xiaomengyang, 850 m a.s.l., 29.iii.1957, 2 1 ♀, L.-C. Zang lgt., P. Kment det. ( NMPC); Tengchong, S margin of the city, 24°59.4′N 98°28.9′E, 1650 m a.s.l., [Ch12], individually collected under stones, on soil surface, on plants and shrubs, in pool with clay bottom, margin of mixed forest above the rice fields, 1 ♀, J. Hájek & J. Růžička lgt., P. Kment det. ( NMPC); Gengma, 5.v.1955, 1 ♀, T.-R. Huang lgt., P. Kment det. ( NMPC); 30 km SW of Jinping, Měnglā, 400 m a.s.l., 3.v.1956, 1 ♀, K.-R. Huang lgt., P.Kment det. ( NMPC);Jingdong, 1200 m a.s.l., 21.iii.1957, 1, A. Monchadskiy lgt., P. Kment det. ( NMPC). IRAN: Iran, no more details, 6.ix.1961, 28 18 ♀♀, J. Klapperich lgt., P. Kment det. ( NMPC). ALBORZ: Kandavan – pass, S slope, 2700–2900 m a.s.l., (36°10′N 51°15′E, Loc. No. 395), 4.–9. vii.1977, 1 ♀, Exp. Nat. Mus. Praha lgt., P. Kment det. ( NMPC); Karaj, 1200 m a.s.l., 23.vi.1960, 3, J. Klapperich lgt., P.Kment det. ( NMPC); Karaj, 3 km N of Dam, 1650 m a.s.l. (35°48′N 50°59′E, Loc. No. 85), 10.viii.1970, 2, Exp. Nat. Mus. Praha lgt., P. Kment det. ( NMPC). ARDABIL: 5 km NW Kolur, ʻShahrud’ river valley, 37°26.1′N 48°41.2′E, 1670 m a.s.l., 4.–5.vi.2006, 1 ♀, J. Hájek & P. Chvojka lgt., P. Kment det. ( NMPC). FARS: 5 km E of Furk, on the road Hajiabad–Darab, 900 m a.s.l. (28°18′N 55°16′E, Loc. No. 217), 28.v.1973, 1 ♀, Exp. Nat. Mus. Praha lgt., P. Kment det.( NMPC);Aliabad, 75 km NW Jahrom (29°30′N 52°35′E; Loc. No. 53), 10.vii.1970, 1 1 L, Exp. Nat. Mus. Praha lgt., P. Kment det. ( NMPC); Dasht-e Arzhan, 50 km W of Shiraz, Shur river (29°39′N 51°58′E, Loc. No. 44), 5.vii.1970, 2 2 L, Exp. Nat. Mus. Praha lgt., P. Kment det. ( NMPC); Tang-e Chogan-e Olia, ruins of Bishapur, 1000 m a.s.l. (29°47′N 51°36′E, Loc. No. 233), 10.vi.1973, 4

1 L, Exp. Nat. Mus. Praha lgt., P. Kment det. ( NMPC). GILAN: Racht [= Rasht], 7.v.1965, 1 1 ♀, Safavi lgt., P. Kment det. ( NMPC). HORMOZ- GAN: 20 km SE of Minab (27°05′N 57°14′E, Loc. No. 204), 21.v.1973, 1 ♀, Exp. Nat. Mus. Praha lgt., P. Kment det. ( NMPC); Bashagerd [mts.], Senderk, 11.v.1977, 1 ♀, Safavi & Pazuki lgt., P. Kment det. ( NMPC); Faryab, Rud-e Rudan river, 350 m a.s.l. (27°28′N 57°07′E, Loc.No.201), 17.–18.v.1973, 1 ♀, Exp. Nat. Mus. Praha lgt., P. Kment det. ( NMPC); Teleng, 40 km SE of Minab (26°53′N 57°16′E, Loc. No. 205), 21.v.1973, 1 2 ♀♀ 2 L, Exp. Nat. Mus.Praha lgt., P.Kment det. ( NMPC). KERMAN: 33 km W Sabzvaran [= Jiroft] (28°44′N 57°28′E, Locality No. 189), 1100 m a.s.l., 6.–7.v.1973, 1 ♀, Exp. Nat. Mus. Praha, P. Kment det. ( NMPC); Deh Bakri (29°03′N 57°56′E, Locality No. 186), 1700–1750 m a.s.l., 30.iv.–3.v. 1973, 1 1 ♀ 1 spec., Exp. Nat. Mus. Praha lgt., P. Kment det.( NMPC). QAZVIN: Hasan Sabbah Castle, 36°27.0′N 50°34.9′E, 2075 m a.s.l., streamlet valley, 13.v.2008, 1 ♀, J. Hájek & P. Chvojka lgt., P. Kment det.( NMPC). SISTAN AND BALUCHISTAN: 16 km SE of Tangé-Sarhé, 61 km NNW Nikshahr, 900 m a.s.l. (26°29′N 60°02′E, Loc. No. 154), 10.iv.1973, 3 ♀♀, Exp. Nat. Mus. Praha lgt., P. Kment det. ( NMPC); Taftan [Mts.], Tamandan Valley (28°36′N 61°02′E, Locality No. 167), 2100 m a.s.l., 20.iv.1973, 1 ♀, Exp. Nat. Mus. Praha lgt., P. Kment det. ( NMPC). TEHRAN: Elburgsgeb. [= Elburz Mts.], Laschkarak Tal [= Lashgarak valley], 1800 m a.s.l., vii.–x. 1961, 1 3 ♀♀, J. Klapperich lgt., P. Kment det. ( NMPC). ZANJAN: 7 km E Sa’id Abad, 36°23.3′N 48°11.7′E, 1635 m a.s.l., 11.–12.v.2008, steppe stream, at light, 1 ♀, J. Hájek & P. Chvojka lgt., P. Kment det. ( NMPC). OMAN: NE, 17 km W of Sur, 15.iii.2015, 1 1 ♀, M. Snížek lgt., P. Kment det. ( ZJPC). TUR- KEY: ADIYAMAN: Burmapinar env., bank of river in canyon above Septimus Severus bridge, 37°55′57″ 38°36′31″, 704 m a.s.l., 3.vii.2011, 2 1 L1 1 L2, P.Kment lgt. & det. ( NMPC). ANTALYA: Köprülü canyon, Beşkonak, 80 km NE of Antalya, 7.ix.1992, 3 1 L, Z. Jindra lgt., P. Kment det. ( ZJPC). TUNCELI: Ovacik env., bank of Munzur river downstreams the village before entering Munzur Vadisi Milli Parki, 4.vii.2011, 7 4 ♀♀ 1 L, P. Kment lgt. (6 4 ♀♀ NMPC, 1 1 L JDKD). Afrotropical Region: ANGOLA: Angola (No. 10238.12), 1 ♀, P. Kment det. ( NMPC). LUNDA NORTE: Luachimo (Ang. No. 915.6), 11.vii.1948, 1 1 ♀, A. B. Machado lgt., P. Kment det. ( NMPC); Luachimo (M) (Ang. No. 3967.2), 6.vi.1959, 1 2 ♀♀, Mus. do Dundo, P. Kment det. ( NMPC); Luachimo, Route Tourismo, M. H. B (Ang. No. 20175), 28.vi.1957, 1 ♀, Mus. do Dundo, P. Kment det. ( NMPC). LUNDA SUL: Alto Chicapa, Camutongola river, Luhembe (M) (Ang. No. 4174.7), 1, 22.vi.1954, Mus. do Dundo, P. Kment det. ( NMPC); Alto Chicapa, Camutongola river, Luhembe (M) (Ang. No. 4166.10, 4168.7, 4181), 11 11 ♀♀, 24.vi.1954, Mus. do Dundo, P. Kment det. ( NMPC); Alto Chicapa, Tehissango / Cuango Muquè (Ang. No. 4308.14), 1 ♀, 16.vii.1954, Mus. do Dundo, P. Kment det. ( NMPC); Alto Cuílo (M), Cavuemba (Ang. No. 4082.24), 31.v.1954, 3 ♀♀, Mus. do Dundo, P. Kment det. ( NMPC). CENTRAL AFRICAN REPUBLIC: 90 km NNE Bangui, 5°03′N 18°47′E, 380 m a.s.l., 14.iii.2010, 1, J. Halada lgt., P. Kment det.( MMBC). DEMOCRATIC REPUBLIC OF THE CONGO: KASAI OCCIDENTAL: Tshikapa (Ang. No. 880.4), vi.–vii. 1948, 2 1 ♀, A.B. Machado lgt., P. Kment det.( NMPC). NIGERIA: TARABA: Gashaka Gumti NP, 30 km SE Serti env., 7°21′N 11°32′E, ca. 450 m a.s.l., 24.iv. –8.v. 2011, 1 1 ♀, J. Halada lgt., P. Kment det. ( ZJPC). SUDAN: SENNAR: Blue Nile prov., Dinder Game Reserve, Galegu, 4.iii.1967, 1

2 ♀♀, P. Štys lgt. (686), P. Kment det. ( MMBC). Oriental Region: IN- DIA: MAHARASHTRA: 10 km E from Savantvadi, 22.v.2006, 2, Z. Kejval lgt., P. Kment det. ( MMBC); 15 km E of Savantvadi, 15°55′N 75°53′E, 40 m a.s.l., riverside, 22.v.2006, 1, Z. Kejval lgt., P. Kment det. ( MMBC); 40 km W of Pune, Mulshi env., 30.ix.–2.x.2005, 1 ♀, J. Bezděk lgt., P.Kment det.( NMPC); Western Ghats Mts., Panchgani, Wai env., 3.–5.vi.2008, 2 1 ♀, Z. Kejval lgt., P. Kment det. ( MMBC). PUDUCHERRY: Karaikal, Kurumbagaram, 23.iii.1947, 48 25 ♀♀, Nathan lgt., L. Hoberlandt det., P. Kment revid. (43 24 ♀♀ NMPC, 5 1 ♀ MMBC). RAJASTHAN: 20 km N of Dausa, Golakabas env., Sarsadevi, 27°05.46″N 78°17.18″E [sic!], 380 m a.s.l., 19.xi.2002, P. Šípek lgt., P. Kment det. ( NMPC); 20 km N of Dausa, Naranimata env., 27°05.46″N 78°17.18″E [sic!], 400 m a.s.l., light trap, 7.–8.xi.2002, 1 ♀, P. Šípek lgt., P. Kment det. ( NMPC); 50 km W of Agra, Bharatpur env., 27°12.42′N 77°30.48″E, 220 m a.s.l., 31.viii.–5.ix.2002, 1 ♀ 1 L5, P. Šípek & M. Fikáček lgt., P. Kment det. ( NMPC). TAMIL NADU: Madras occ., Tiruchirappalli, 16.v.1970, 1 ♀, J. Hloušek lgt., P. Kment det. ( MMBC). LAOS: VIENTIANE: Phou Khao Khouay NBCA, ca. 50 km N Vientiane (lake env.), 18°23.2′N 102°45.5′E, 745 m a.s.l., 18.v.2010, 1, J. Hájek lgt., P. Kment det. ( NMPC); Vang-Veng, 18°55′23″N 102°26′53″, 300 m a.s.l., 10.–15.v. & 1.–6.vii.2001, 1, J. Kolibáč lgt., P. Kment det. ( MMBC). MALAYSIA: SABAH: Mt. Kinabalu env., 28.v.1999, M. Snížek lgt., P. Kment det. ( ZJPC); Sapulut env., 1.–6. vi.2001, 1 ♀, J. Stolarczyk lgt., P. Kment det. ( ZJPC); W, Crocker Range E, W of Apin Apin, v.1999, 1, M. Snížek lgt., P. Kment det. ( ZJPC). SARAWAK: Kapit distr., Rumah Ugap env., Sut river valley, 3.–9.iii.1994, 2 ♀♀, P. Bílek lgt., P. Kment det. ( ZJPC); Kapit distr., Sebong, Baleh river, 6.–21.iii.1994, 2, S. Bílý lgt., P. Kment det. ( NMPC). PAKIS- TAN: PUNJAB: Rawalpindi env., Basal, Kalachitta Range, 4.–7.i.1956, 1

, Chr. Lindemann lgt., L. Hoberlandt det., P. Kment revid. ( NMPC). SRI LANKA: CENTRAL PROVINCE: Sigirya near Dambulla, Pidurangala archeological areal, 7°58.18′N 80°45.60′E, 24.ii., 5 2 ♀♀ (4 1 ♀ ZJPC, 1 1 ♀ NMPC). RATNAPURA: Panamure env., 11.xii.1995, 1 ♀, S. Bečvář lgt., P. Kment det. ( MMBC). SOUTHERN PROVINCE: Tissamaharama env., 90 km NE of Matara, 14.iii.1994, 1 1 ♀, Z. Kejval lgt., P. Kment det. ( ZJPC). THAILAND: CHIANG MAI: Doi Suthep Mt., 1.v.1996, 1, S. Bílý lgt., P. Kment det. ( NMPC). SATUN: Thale Ban, 6°45′N 100°09′E, 200 m a.s.l., 8.–13.iv.1997, 1 ♀, J. Kolibáč lgt., P. Kment det. ( MMBC).

Distribution. Palaearctic Region: Europe: Bulgaria (JOSI- FOV 1954, 1960, 1963; GÖLLNER- SCHEIDING & ARNOLD 1988; STOIANOVA & SIMOV 2016); France ( LATREILLE 1804, 1807; DUFOUR 1833; BURMEISTER 1835; BRULLÉ 1836; AMYOT & SERVILLE 1843; PUTON 1880; POISSON 1925, 1957; DE SEABRA 1926, JACZEWSKI 1934; RIEGER 1976, 1977; ELDER 2017); Gibraltar ( MAYR 1868, HOFFMANN 1941); Greece ( JOSIFOV 1959, ZIMMERMANN 1982, POLHEMUS & POLHEMUS 2012); Italy, incl. Sicily ( COSTA 1862, MANCINI 1952, SERVADEI 1967, TAMANINI 1981, CIANFERONI & TERZANI 2013); North Macedonia ( PROTIĆ 1998); Portugal (DE SEABRA 1926, NIE- SER & MONTES 1984, BAENA & VÁZQUEZ 1986, HEBSGAARD et al. 2004); Russia (Daghestan) ( KANYUKOVA 1997, 2006); Spain ( DUFOUR 1833; BURMEISTER 1835, DE SEABRA 1926, LINDBERG 1929, BAENA RUIZ & FERRERAS ROMERO 1982, NIESER & MONTES 1984, BAENA & VÁZQUEZ 1986, GARCÍA ROJAS et al. 1987, NIESER & CHEN 1992, PAGOLA- CARTE & RIBES 2007, GÜNTHER & GÜNTHER 2019); Turkey: European Part ( FENT et al. 2011, FENT & DURSUN 2018). North Africa: Algeria ( SIENKIEWICZ 1964, ECKERLEIN & WAGNER 1965); Egypt ( PRIESNER & ALFIERI 1953, LINNAVUORI 1964); Morocco ( LINDBERG 1929, VIDAL 1937, AGUESSE et al. 1982, GHEIT 1995, L’MOHDI et al. 2008); Tunisia ( LINDBERG 1922, CARAPEZZA 1997, SLIMANI et al. 2015). Asia: Afghanistan ( CHINA & MILLER 1950); Armenia ( AKRAMOVSKAYA 1959, KANYUKOVA 2006); Azerbaijan, incl. Nakhichevan Republic ( KOLENATI 1857; HORVÁTH 1899; KIRITSHENKO 1918, 1938; KANYUKOVA 2006); China: Anhui (new record), Beijing ( NIESER & CHEN 1992, CHEN 1999, XIE & LIU 2013), Fujian ( KORMILEV 1971, NIESER & CHEN 1992, CHEN 1999, HUA 2000, XIE & LIU 2013), Guangdong ( HOFFMANN 1933, 1941; NIESER & CHEN 1992; CHEN 1999; HUA 2000; CHEN et al. 2001; XIE & LIU 2013), Guizhou ( NIESER & CHEN 1992, CHEN 1999, XIE & LIU 2013), Hainan ( HOFFMANN 1933, 1941; NIESER & CHEN 1992; CHEN 1999; XIE & LIU 2013), Hebei ( LIU & BU 2009), Henan ( SHEN et al. 2014, CAI et al. 2016), Hong Kong ( HOFFMANN 1941, KORMILEV 1971), Hubei ( NIESER & CHEN 1992, CHEN 1999, XIE & LIU 2013), Hunan ( NIESER & CHEN 1992, CHEN 1999, XIE & LIU 2013), Inner Mongolia ( NIESER & CHEN 1992, CHEN 1999, XIE & LIU 2013), Jiangsu ( NIESER & CHEN 1992, XIE & LIU 2013), Jiangxi ( ZHANG 1994, HUA 2000, LI et al. 2008), Shanxi ( MIYAMOTO 1960, as O. m. flavomarginatus), Shaanxi (new record), Sichuan ( NIESER & CHEN 1992, CHEN 1999, HUA 2000, XIE & LIU 2013), Tianjin ( NIESER & CHEN 1992, CHEN 1999), Xizang (= Tibet) ( HUA 2000), Yunnan ( CHENG et al. 2006), Zhejiang ( KORMILEV 1971, NIESER & CHEN 1992, CHEN 1999, XIE & LIU 2013); Cyprus ( LINDBERG 1948, GEORGHIOU 1977, POLHEMUS & POLHEMUS 2012); Georgia ( KIRITSHENKO 1918, KANYUKOVA 2006); Iran ( LINNAVUORI & HOSSEINI 2000, GHAHARI et al. 2013); Israel ( POLHEMUS & POLHEMUS 2012); Japan ( SCOTT 1874, as P. flavomarginatus): Hokkaido ( HAYASHI 1995, as P. flavomarginatus; HAYASHI et al. 2016; HAYASHI & MIYAMOTO 2018), Honshu ( MATSUMURA 1905, 1915, as P. flavomarginatus; HIURA 1967, as O. m. flavomarginatus; KORMILEV 1971; TOMO- KUNI 1981, as O. m. flavomarginatus; HAYASHI et al. 2016; HAYASHI & MIYAMOTO 2018), Kyushu ( MATSUMURA 1905, 1915, as P. flavomarginatus; JACZEWSKI 1934; KORMILEV 1971; HIURA 1967, as O. m. flavomarginatus; TAKENO 1998; HAYASHI et al. 2016; HAYASHI & MIYAMOTO 2018), Ryukyu Islands ( MIYAMOTO 1964, HAYASHI et al. 2016; HAYASHI & MIYAMOTO 2018), Shikoku ( HIURA 1967, as O. m. flavomarginatus; YANO et al. 2012; HAYASHI et al. 2016; HAYASHI & MIYAMOTO 2018), Tanegashima Islands ( HAYASHI & MIYAMOTO 2018); Jordan ( KATBEH et al. 2000); Korea ( MIYAMOTO & LEE 1966, JOSIFOV & KERZHNER 1972, LEE & KWON 1991, LEE et al. 1993, LEE et al. 2013); Lebanon ( LINDBERG 1922); Oman (new record); Russia: Far East: Kunashir Island ( KERZHNER 1978, KERZHNER & MARUSIK 1997, KERZHNER et al. 2004, KANYUKOVA 2006, KANYUKOVA & MARUSIK 2006, VINOKUROV et al. 2010); Saudi Arabia ( BROWN 1951, LINNAVUORI 1986); Syria ( FIEBER 1861, POISSON 1934, NIESER & MOUBAYED 1985); Taiwan (MAT - SUMURA 1915, as P. formosanus; BERGROTH 1914; ESAKI 1926; JACZEWSKI 1934, KORMILEV 1971, XIE & LIU 2013); Turkmenistan ( KIRITSHENKO 1957, KANYUKOVA 2006); Turkey (Asian part) ( LINDBERG 1922, HOBERLANDT 1952, FENT et al. 2011, MATOCQ et al. 2014, ÖZGEN et al. 2017, ÇERÇI et al. 2018, DURSUN & FENT 2018, FENT & DURSUN 2018); United Arab Emirates ( LINNAVUORI et al. 2011). Afrotropical Region: Angola ( KORMILEV 1973); Botswana ( KRÜGER & DECKERT 2016); Burkina Faso ( LINNAVUORI 1981); Cameroon ( KORMILEV 1973); Cape Verde Islands ( LINDBERG 1959); Central African Republic (new record), Democratic Republic of the Congo ( KORMILEV 1973); Ethiopia ( JACZEWSKI 1934, BROWN 1951); Gambia ( NIESER & CHEN 1992); Guinea-Bissau ( MANCINI 1939a); Ivory Coast ( LINNAVUORI 1981);? Kenya ( MONTANDON 1914); Mozambique ( POISSON 1934);? Namibia ( HESSE 1925); Nigeria ( KORMILEV 1973, LINNAVUORI 1981);? Senegal ( SIENKIEWICZ 1964); South Sudan ( LINNAVUORI 1980); Sudan (new record); Tanzania ( NIESER & CHEN 1992, POLHEMUS & POLHEMUS 2012);? Uganda ( MONTANDON 1914); Yemen ( BROWN 1951, LINNAVUORI 1989). Oriental Region: India ( DISTANT 1906): Assam (THIRUMALAI 2007, POLHEMUS & POLHEMUS 2012), Bihar (MAXWELL- LEFROY 1909, DISTANT 1910, KATIYAR 1953, THIRUMALAI 2007), Kerala ( DISTANT 1910, THIRUMALAI 2007), Madhya Pradesh (THIRUMALAI 2007, THIRUMALAI et al. 2007), Maharashtra ( PAIVA 1919b; JACZEWSKI 1934; THIRUMALAI & SHARMA 2002, 2012), Manipur ( BAL & BASU 2004, THIRUMALAI 2007), Meghalaya ( PAIVA 1919 a, BAL & BASU 1998, THIRUMALAI 2007), Mizoram ( BAL & BASU 2007), Orissa ( DISTANT 1910, GHOSH et al. 1989, THIRUMALAI 2007), Puducherry: Karaikal (KOR- MILEV 1971, THIRUMALAI 2007), Rajasthan (new record),

Tamil Nadu ( KATIYAR 1953; SIENKIEWICZ 1964; THIRUMALAI 1989, 1999, 2001, 2007; POLHEMUS & POLHEMUS 2012), Tripura ( BAL & BASU 2000), Uttar Pradesh ( DISTANT 1910, THIRUMALAI 2007), West Bengal ( DISTANT 1910, THIRUMA- LAI 2007, POLHEMUS & POLHEMUS 2012); Indonesia: Bali ( SCHOUTEDEN 1933, POLHEMUS & POLHEMUS 2012), Java ( JACZEWSKI 1935, KORMILEV 1971), Sumatra ( JACZEWSKI 1934, MANCINI 1939a); Laos ( KORMILEV 1971); Malaysia: Peninsular part ( KORMILEV 1971, KOVAC & YANG 1989, POLHEMUS & POLHEMUS 2012), Sabah ( KORMILEV 1971; NIESER & CHEN 1992, 1999), Sarawak ( KORMILEV 1971); Myanmar ( DISTANT 1910, JACZEWSKI 1934); Nepal (DIS- TANT 1910); Pakistan ( HAMID & HAMID 1973); Singapore ( POLHEMUS & POLHEMUS 2012); Sri Lanka ( DISTANT 1910, as P. marginatus, 1915 , as S. rutherfordi; JACZEWSKI 1934; POLHEMUS 1979; POLHEMUS & POLHEMUS 2012); Thailand ( KORMILEV 1971, POLHEMUS & POLHEMUS 2012); Vietnam ( DISTANT 1906, JACZEWSKI 1934, as Cochinchine; KORMILEV 1971, POLHEMUS & POLHEMUS 2012). Australasian Region: Indonesia: Sulawesi ( NIESER & CHEN 1999).

Notes. POLHEMUS (1995a) listed O. marginatus from Bosnia and Herzegovina and Croatia, however, there are no such records in the relevant literature (cf. JOSIFOV 1986, PROTIĆ 1998, KMENT & BERAN 2011) and taking into account its confinement to the southernmost regions of Balkan Peninsula ( JOSIFOV 1986, STOIANOVA & SIMOV 2016) they are probably based on misinterpretation of records from former Yugoslavia. The records of O. marginatus from Australia ( KIRKALDY 1908, based on a larva; HALE 1925), Tasmania, New Guinea and New Caledonia ( MONTANDON 1897) belong to other species. The record from New Caledonia ( DISTANT 1920) belong to O. australicus ( JACZEWSKI 1934). The records of O. marginatus from Kenya, Uganda ( MONTANDON 1914), Namibia ( HESSE 1925) or Senegal ( SIENKIEWICZ 1964) requires confirmation. The records from Myanmar by MONTANDON (1897) and DISTANT (1906) refer to O. feae . POLHEMUS & POLHEMUS (2012) discussed variability of Oriental populations of O marginatus , finding the male genitalia of specimens from Malaysia and Singapore structurally similar to those from India and the west Palaearctic Region. The populations from Bali are smaller and have also slightly different shapes of the superior appendage on the right paramere and the process of the male pygophore. They wrote that these more eastern populations may be assignable to the subspecies O. marginatus insularis from Philippines, but an analysis of further material from throughout the Malay Archipelago region will be necessary to properly determine the subspecific designations that should be applied to these insular forms ( POLHEMUS & POLHEMUS 2012). According to N. Nieser and P.-p. Chen, O. marginatus in its present sense represents rather a complex of several species ( CHEN et al. 2005). In this paper we identify O. m. marginatus as a polymorphic species in the sense of JACZEWSKI (1934), KOR- MILEV (1973) and POLHEMUS & POLHEMUS (2012). However, we observed considerable differences in the shape of the parameral cap and the parameral appendages among the populations, e.g when comparing specimens from tropical Africa, Mediterranean and south India, and agree with the above-mentioned opinion of CHEN et al. (2005).

Figures

Figs 4–8. 4–6 – Ochterus papaceki sp. nov.,, Arher:4 – body, dorsal view (magnification 27×); 5 – head, frontal view (80×); 6 – paramere, ventral view (magnification 180×). 7 – O. marginatus (Latreille, 1804), head, frontal view (75×) (, Turkey, Tunceli Prov., Ovacik env., NMPC). 8 – O. caffer (Stål, 1855), head, frontal view (70×) (, Democratic Republic of the Congo, Ituri, NMPC). Scale bars: 0.2 mm (Fig. 6), 0.5 mm (Figs 5, 7–8), 1 mm (Fig. 4).