Manota arenalensis, Jaschhof & Hippa, 2005

Jaschhof, Mathias & Hippa, Heikki, 2005, The genus Manota in Costa Rica (Diptera: Mycetophilidae), Zootaxa 1011 (1), pp. 1-54 : 25-27

Manota arenalensis sp. n. ( Fig. 12 )

Description. Large species.

Male. Head. Antenna with fourth flagellomere 1.9 times as long as wide. Maxillary palpus with third palpomere bearing 3 curved sensilla; fourth palpomere with a parasegment apically.

Thorax. Anterior basalare and laterotergite each bare. Episternum 3 with ventral portion setose. Wing. Length 1.7–2.2 mm. Membrane with a few setae along posterior margin. Sc bare. Junction of CuA2 into CuA1 incomplete or complete.

Terminalia ( Fig. 12 ). Sternite 9 almost completely merged with gonocoxites, with only small portions of posterior margin recognisable; margin situated beyond mid length of gonocoxites. Ventro­lateral edges of gonocoxites lobe­like and with some 5 stronger setae; interiorly with 1 curved seta on a separate rim. Disto­lateral lobes of gonocoxites large; with large, straight setae pointing posteriorly and smaller, curly setae pointing inwards. Gonocoxites in position I with 1 medium­sized, blunt megaseta; in position II with 3 short, strong, stiff setae on lobe; in position III with 2 flat megasetae, one megaseta simple and pointed, the other two­branched with one branch broadly rounded and the other branch whip­like; in position IV with 1 pointed megaseta directed inwards. Gonostyli small, slender, bilobed; ventral lobe each with 1 large and 1 smaller setae apically, dorsal lobe with 2+1 setae with one apical seta strongly re­curved. Parastylar lobes large, very weak in some specimens, but seemingly sclerotised in others. Tegmen long and narrow, almost evenly tapering to tip. Hypoproct broad apically; with long setae laterally.

Diagnostic characters. Among the species with both anterior basalare and laterotergites bare, it has the terminalia characterised by the combination of very small, slender gonostyli; gonocoxites in position IV with one large, pointed megaseta pointing strictly inwards (see also next species, M. eximia ); and sternite 9 almost indistinguishable from the gonocoxites.

Etymology. Derived from the type locality, Arenal volcano.

Types. Holotype. Male. Costa Rica, Alajuela province, San Carlos, Volcán Arenal National Park, Sendero Pilón , 650 m, 6–7 Nov. 2000, by Malaise trap, G. Carballo ( INBio sample #60084) . Paratypes. 3 males, same data as the holotype ; 3 males, Cartago province, Tapantí­Macizo de la Muerte National Park, Sector la Represa , 1550 m, 26 March 2000, by Malaise trap, D. Rubí & C. Hansson ( INBio sample #67894) ; 2 males, same National Park, but Sendero Arboles Caídos , 1500–1600 m, 9 April 2002, by Malaise trap, D. Rubí & C. Hansson ( INBio sample #67891) .

Other material studied. 1 male, Puntarenas province, La Amistad International Park, Buenos Aires, Sendero a Casa Coca , 13 Oct.–13 Nov. 2001, D. Rubí ( INBio sample #74889) ; 3 males, Limón province, Hitoy Cerere Biological Reserve, Sendero Espavel , 18 Sep.–7 Oct. 2003, E. Rojas, B. Gamboa, W. Arana, M. & C. Jaschhof ; 1 male, Limón province, Pococí, Braulio Carrillo National Park, Estación Quebrada González , 21 Aug. 2002, P. Hanson & C. Godoy ( INBio sample #70777) .

Figures

FIGURE 12. Manota arenalensis sp. n. A: Male terminalia and sternite 8, ventral view (holotype). B: Male terminalia, dorsal view (paratype). C: gonostylus, ventral view (paratype from Tapantí). Scale = 0.1 mm.