Manota acuminata, Jaschhof & Hippa, 2005

Jaschhof, Mathias & Hippa, Heikki, 2005, The genus Manota in Costa Rica (Diptera: Mycetophilidae), Zootaxa 1011 (1), pp. 1-54 : 36-37

Manota acuminata sp. n. ( Fig. 18 )

Description. Small species.

Male. Head. Antenna with fourth flagellomere 1.7 times as long as wide. Maxillary palpus with third palpomere bearing 3 curved sensilla; fourth palpomere with a parasegment apically.

Thorax. Anterior basalare and laterotergite each bare. Episternum 3 setose. Wing. Length 1.5–1.6 mm. Membrane with a few setae along posterior margin. Sc bare, or with 1 single seta dorsally. Junction of CuA2 into CuA1 incomplete.

Terminalia ( Fig. 18 ). Sternite 9 with only posterior margin distinct; margin situated at mid length of gonocoxites, strongly emarginate. Ventro­lateral lobes of gonocoxites subrectangular; interiorly each with 1 long seta on a separate rim. Disto­lateral lobes of gonocoxites short, bilobed; outer lobe with very large, straight setae; inner lobe subcircular, with long, curly setae. Gonocoxites in position I+II with 7–9 short, blunt megasetae; in position III with 2 flat megasetae with one megaseta whip­like and the other rounded apically; in position IV with 2 long, pointed megasetae directed inwards. Gonostyli small, flat; each bilobed posteriorly, one lobe bare, the other lobe bearing 1 strongly re­curved seta apically; antero­apically with 1 large, straight seta directed inwards plus 1 short, fine seta. Parastylar lobes largely hidden behind gonostyli, large, bare, almost pointed posteriorly (not illustrated). Tegmen wide basally, strongly narrowed in distal half. Hypoproct broad apically, with posterior margin of tergite 10 behind; with a few rather long setae laterally.

Diagnostic characters. Among the species with both anterior basalare and laterotergites bare, and terminalia with sternite 9 barely discernible, it has the disto­lateral lobes of the gonocoxites large, and the gonostyli small and flat. The array of megasetae dorsally on the gonocoxites is species specific (in particular, the conspicuous megasetae in positions I+II and IV), as is the large, bare, pointed parastylar lobe.

Etymology. From Latin, acuminatus, pointed, referring to the outline of the parastylar lobe.

Types. Holotype. Male. Costa Rica, Alajuela province, San Carlos, Volcán Arenal National Park, Sendero Pilón , 650 m, 6–7 Nov. 2000, by Malaise trap, G. Carballo ( INBio sample #60084) . Paratypes. 5 males, same data as the holotype ; 2 males, Limón province, Hitoy Cerere Biological Reserve, Sendero Espavel , 560 m, 11 March–1 April 2003, by Malaise trap, E. Rojas, B. Gamboa & W. Arana ( INBio sample #73628) .

Other material studied. 1 male, Puntarenas province, Corcovado National Park , 600 m S Cerro Rincón, 23 April–24 June 2002, J. Azofeifa ( INBio sample #70842) ; 2 males, same locality, but 1 Aug.–25 Sep. 2002 ( INBio sample #71459) ; 1 male, Limón province, Pococí, Braulio Carrillo National Park, Estación Quebrada González , 21 Aug. 2002, P. Hanson & C. Godoy ( INBio sample #70777) ; 1 male, Limón province, Hitoy Cerere Biological Reserve, Sendero Espavel , 18 Sep.–7 Oct. 2003, E. Rojas, B. Gamboa, W. Arana, M. & C. Jaschhof ; 1 male, Cartago province, Tapantí–Macizo de la Muerte National Park, Estación La Esperanza , 3–22 Nov. 2003, M. & C. Jaschhof .

Figures

FIGURE 18. Manota acuminata sp. n. A: Male terminalia, ventral view (holotype). B: Male terminalia, dorsal view (paratype from Hitoy Cerere). Scale = 0.1 mm.