PSEUDOCOCCIDAE

Franco, José Carlos, Russo, Agatino & Marotta, Salvatore, 2011, An annotated checklist of scale insects (Hemiptera: Coccoidea) of Portugal, including Madeira and Azores Archipelagos, Zootaxa 3004, pp. 1-32 : 18-21

FAMILY PSEUDOCOCCIDAE

Atrococcus paludinus (Green, 1921)

Distribution: Mainland

References: Kozár & Franco ( 1995), Ben-Dov et al. ( 2010), Danzig & Watson ( 2010).

Balanococcus scirpi (Green, 1921)

Distribution: Mainland

References: Kozár & Franco ( 1995), Ben-Dov et al. ( 2010), Danzig & Watson ( 2010).

Cucullococcus arrabidensis (Neves, 1954)

Distribution: Mainland

References: Neves ( 1954 b, 1956), Ben-Dov ( 1994), Marotta & Franco ( 1999), Ben-Dov et al. ( 2010), Danzig & Watson ( 2010).

● Delottococcus proteae (Hall, 1937)

Distribution: Madeira

Comments: first record; collected and identified by A.M.F. Aguiar on Protea magnific a x susannae (var. Suzara), in Santo da Serra (Santa Cruz) ( 19 - 10-2000); on Protea cynaroides , in Caniço (Santa Cruz) ( 7 -09- 2000) and Santana ( 22 -08- 2006); on Leucadendron tinctum , Protea sp., Protea repens , Protea longifolia “Satin Pink”, in Santa ( 19 - 10-2000).

Dysmicoccus boninsis (Kuwana, 1909)

Distribution: Madeira

References: Green ( 1923), Balachowsky ( 1939), Williams ( 1970), Vieira et al. ( 1983), Ben-Dov ( 1994), CABI ( 1995 b), Ben-Dov et al. ( 2010), Borges et al. ( 2008), Danzig & Watson ( 2010), Pellizzari & Germain ( 2010).

Dysmicoccus brevipes (Cockerell, 1893)

Distribution: Azores and Madeira

References: Bensaúde ( 1926 a), Balachowsky ( 1939), Borchsenius ( 1949), CAB ( 1972 c), Vieira et al. ( 1983), Ben-Dov ( 1994), Borges ( 2005), Ben-Dov et al. ( 2010), Borges et al. ( 2008), Danzig & Watson ( 2010), Pellizzari & Germain ( 2010).

Dysmicoccus grassii (Leonardi, 1913)

Ditribution: Madeira

References: Williams & Granara de Willink ( 1992).

Comments: Williams & Granara de Willink ( 1992) reported (pp. 13) the presence of this species as D. alazon .

Nipaecoccus nipae (Maskell, 1893)

Distribution: Mainland, Azores and Madeira.

References: Bensaúde ( 1926 b), Neves ( 1935, 1936c), Balachowsky ( 1939), Seabra ( 1942), CAB ( 1966 b), Guimarães ( 1973), Carneiro ( 1982), Fernandes ( 1982), Vieira et al. ( 1983), Ben-Dov ( 1994), Carvalho & Aguiar ( 1997), Carvalho et al. ( 1997), Carvalho ( 1999), Borges ( 2005), Ben-Dov et al. ( 2010), Franco et al. ( 2006), Borges et al. ( 2008), Danzig & Watson ( 2010), Pellizzari & Germain ( 2010).

● Paracoccus leucadendri Mazzeo & Franco

Distribution: Mainland and Madeira

References: Mazzeo et al. ( 2009), Ben-Dov et al. ( 2010).

Comments: although described from specimens collected in Portugal, it is possibly native to Southern Africa ( Mazzeo et al., 2009); new record for Madeira, collected and identified by A.M.F. Aguiar on Leucadendron sp. in Ribeirinha (Camacha) ( 7 -05- 1997).

Phenacoccus latipes Green, 1923

Distribution: Madeira

References: Green ( 1923), Balachowsky ( 1939), Vieira et al. ( 1983), Ben-Dov ( 1994), Ben-Dov et al. ( 2010), Borges et al. ( 2008), Danzig & Watson ( 2010).

Phenacoccus madeirensis Green, 1923

Distribution: Mainland and Madeira

References: Green ( 1923), Balachowsky ( 1939), Vieira et al. ( 1983), Ben-Dov ( 1994), Carvalho ( 1999), CABI ( 2000), Ben-Dov et al. ( 2010), Borges et al. ( 2008), Danzig & Watson ( 2010), Pellizzari & Germain ( 2010).

Comments: the presence of this mealybug in Mainland reported by Danzig & Watson ( 2010) is confirmed by us: specimens collected by Elisabete Figueiredo on Capsicum annuum under greenhouse, in Silveira (Torres Vedras) ( 30 - 7-2009).

Phenacoccus monieri Balachowsky, 1939

Distribution: Madeira

References: Balachowsky ( 1939), Vieira et al. ( 1983), Ben-Dov ( 1994), Ben-Dov et al. ( 2010), Borges et al. ( 2008), Danzig & Watson ( 2010).

● Phenacoccus peruvianus Granara de Willink, 2007

Distribution: Mainland and Madeira

References: Beltrà et al. ( 2010)

Comments: first record for Madeira; collected on Bougainvillea sp. in Funchal, by A.M.F. Aguiar ( 29 -05- 2001; 12 -06- 2002), by the Phytossanitary Services ( 2 -07- 2001), and by A. Azevedo ( 30 -08- 2001); in Câmara de Lobos, by A.M.F. Aguiar ( 22 -08- 2002), and by V. Balmés ( 10-11 - 2010); and in Porto Santo, by A.M.F. Aguiar ( 25 -07- 2001). All specimens were identified by A.M.F. Aguiar, except those collected in 2010, whose identification was carried out by J.-F. Germain.

● Phenacoccus yerushalmi Ben-Dov, 1985

Distribution: Mainland

Comments: first record; collected by Luis Silva on Pinus sp. in Costa da Caparica ( 3 -06- 1995).

Planococcus citri (Risso, 1813)

Distribution: Mainland, Azores and Madeira

References: Almeida ( 1898 b), Lindinger ( 1912), Green ( 1923), Bensaúde ( 1924, 1926c, 1927 a, 1927 b), Seabra ( 1906, 1926, 1930 a, 1930 b, 1930 c, 1941), Almeida ( 1934, 1935), Neves ( 1936 c, 1952, 1954 a), Balachowsky ( 1939), Borchsenius ( 1949), Guimarães ( 1973), Carneiro ( 1982), Fernandes ( 1982, 1992), Vieira et al. ( 1983), Franco & Carvalho ( 1990), Schanderl et al. ( 1995), Carvalho et al. ( 1997), Franco & Marotta ( 1999), CABI ( 1999 b), Carvalho ( 1994, 1999), Borges ( 2005), Ben-Dov et al. ( 2010), Franco et al. ( 2000, 2006), Borges et al. ( 2008), Danzig & Watson ( 2010), Pellizzari & Germain ( 2010).

● Planococcus ficus (Signoret, 1875)

Distribution: Mainland, Azores and Madeira

References: Almeida ( 1898 b), Seabra ( 1906, 1926, 1941), Almeida ( 1934, 1935), Ezzat & McConnell ( 1956), Carneiro ( 1982), Franco et al. ( 2000), Borges ( 2005), Ben-Dov et al. ( 2010), Danzig & Watson ( 2010). Comments: first record for Madeira; collected by António Miguel Franquinho Aguiar, on Vitis vinifera, in Encruzilhada (S. Vicente) ( 13 - 11-1995).

* Planococcus kraunhiae (Kuwana, 1902)

Distribution: Madeira

References: Balachowsky ( 1939), Vieira et al. ( 1983).

Comments: Balachowsky ( 1939) collected this species on Ficus carica and reported that, apparently, it was not differentiable from Pseudococcus citri Risso v. coleorum Marchal. Vieira et al. ( 1983) did not find it in the Island. This is possibly a misidentification of Planococcus citri (Risso) .

Planococcus minor (Maskell, 1897)

Distribution: Azores and Madeira

References: Borges ( 2005), Borges et al. ( 2008).

● Planococcus vovae (Nasonov, 1909)

Distribution: Mainland

Comments: first record; collected by the first author, on Cupressus sp., in Monsanto (Lisbon) ( 2-10 - 1995, 8- 11-1995) and Setúbal ( 29 -05- 1995) and by Filomena Caetano, on Chamaecyparis sp., in Restelo (Lisbon) ( 6- 11 - 1995).

Pseudococcus calceolariae (Maskell, 1879)

Distribution: Mainland, Azores and Madeira

References: Franco & Carvalho ( 1990), Carvalho ( 1994), Carvalho et al. ( 1997), Franco & Marotta ( 1999), Franco et al. ( 2000, 2006), Ben-Dov et al. ( 2010), Borges et al. ( 2008), Danzig & Watson ( 2010), Pellizzari & Germain ( 2010).

Pseudococcus cimensis Green, 1923

Distribution: Madeira

References: Green ( 1923), Vieira et al. ( 1983), Ben-Dov ( 1994), Ben-Dov et al. ( 2010), Borges et al. ( 2008), Danzig & Watson ( 2010).

Pseudococcus comstocki (Kuwana, 1902)

Distribution: Madeira

References: Balachowsky ( 1939), Borchsenius ( 1949), Vieira et al. ( 1983), Ferreira & Guimarães ( 1984), Ben-Dov ( 1994), Ben-Dov et al. ( 2010), Borges et al. ( 2008), Danzig & Watson ( 2010), Pellizzari & Germain ( 2010).

Pseudococcus longispinus (Targioni Tozzetti, 1867)

Distribution: Mainland, Azores and Madeira

References: Almeida ( 1898 b), Seabra ( 1906, 1930a, 1930 c, 1941), Lindinger ( 1912), Green ( 1923), Bensaúde ( 1927 b), Neves ( 1936 c, 1952, 1954 a), Balachowsky ( 1939), Borchsenius ( 1949), CAB ( 1958 b), Guimarães ( 1973), Carneiro ( 1982), Fernandes ( 1982, 1992), Vieira et al. ( 1983), Franco & Carvalho ( 1990), Ben-Dov ( 1994), Carvalho ( 1994, 1999), Kozár & Franco ( 1995), Schanderl et al. ( 1995), Carvalho & Aguiar ( 1997), Carvalho et al. ( 1997), Franco & Marotta ( 1999), Franco et al. ( 2000, 2006), Borges ( 2005), Ben-Dov et al. ( 2010), Borges et al. ( 2008), Danzig & Watson ( 2010), Pellizzari & Germain ( 2010).

* Pseudococcus maritimus (Ehrhorn, 1900)

Distribution: Madeira

References: Balachowsky ( 1939), Vieira et al. ( 1983)

Comments: reported by Balachowsky ( 1939) from specimens collected at Funchal, Câmara de Lobos and S.Vicente, on several ornamental plants and Trifolium sp., in 1936. According to the present knowledge on the Pseudococcus maritimus complex ( Miller et al. 1984, Gimpel & Miller 1996) this is possibly a misidentification of P. viburni .

Pseudococcus viburni (Signoret, 1875)

Distribution: Mainland, Azores and Madeira

References: Miller et al. ( 1984), Franco & Carvalho ( 1990), Fernandes ( 1993), Carvalho ( 1994, 1999), Kozár & Franco ( 1995), Gimpel & Miller ( 1996), Carvalho et al. ( 1997), Franco & Marotta ( 1999), Franco et al. ( 2000, 2006), Borges ( 2005), Ben-Dov et al. ( 2010), Borges et al. ( 2008), Danzig & Watson ( 2010), Pellizzari & Germain ( 2010).

Rhizoecus martini Kozár & Konczné Benedicty, 2007 Distribution: Madeira

References: Kozár & Konczné Benedicty ( 2007), Ben-Dov et al. ( 2010).

Ripersiella sasae (Takagi & Kawai, 1971) Distribution: Madeira

References: Kozár & Konczné Benedicty ( 2007).

Saccharicoccus sacchari (Cockerell, 1895)

Distribution: Madeira

References: Green ( 1923), CAB ( 1977 b), Vieira et al. ( 1983), Ben-Dov ( 1994), Ben-Dov et al. ( 2010), Borges et al. ( 2008), Danzig & Watson ( 2010).

Seabrina cistorum Neves, 1943

Distribution: Mainland

References: Neves ( 1943), Ben-Dov ( 1994), Ben-Dov et al. ( 2010), Danzig & Watson ( 2010).

Trionymus perrisii (Signoret, 1875)

Distribution: Mainland

References: Kozár & Franco ( 1995), Ben-Dov et al. ( 2010), Danzig & Watson ( 2010).

Vryburgia amaryllidis (Bouché, 1837)

Distribution: Mainland

References: Kozár & Franco ( 1995), Ben-Dov et al. ( 2010), Danzig & Watson ( 2010).