Exocelina cyclops Shaverdo & Balke

Shaverdo, Helena, Sagata, Katayo & Balke, Michael, 2018, Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 803, pp. 7-70: 15-16

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Exocelina cyclops Shaverdo & Balke

sp. n.

5. Exocelina cyclops Shaverdo & Balke  sp. n. Figs 12, 36

Exocelina  undescribed sp. MB3330: Toussaint et al. 2014: supplementary figs 1-4, table 2; Toussaint et al. 2015: supplementary figs S1-S2, table S3.

Exocelina cyclops  New Guinea MB3330: Toussaint et al. 2015: supplementary information S5-S6.

Type locality.

Papua: Jayapura Regency, Cyclops Mts, 02°32.03'S, 140°30.41'E, 710 m a.s.l.

Type material.

Holotype: male "Indonesia: Papua, Cyclops Mts., 710 m, 02°32.031'S, 140°30.412'E, local collector ca. 1997" ( ZSM). Paratypes: 1 male with the same label as the holotype ( NHMW). 1 male, 2 females "Indonesia: Papua, Cyclops Mts., 1120 m, 02°31.516'S, 140°30.436'E, local collector ca. 1997" ( MZB, ZSM). 2 males, 1 female "Indonesia: Papua, Cyclops Mts., 615 m, 02°32.031'S, 140°30.412'E, local collector ca. 1997" ( NHMW, ZSM). 2 males "Indonesia: Papua, Cyclops Mts., Doyo, 365 m, local collector ca. 1997" ( ZSM).


Body size and form: Beetle small: TL-H 3.0-3.25 mm, TL 3.25-3.55 mm, MW 1.65-1.8 mm (holotype: TL-H 3.1 mm, TL 3.4 mm, MW 1.7 mm), with oblong-oval habitus.

Coloration: Reddish. Dorsal surface almost uniformly yellowish red to reddish brown, with paler anterior part of head and pronotum laterally; head appendages and legs yellowish red (Fig. 12). All type specimens are teneral, therefore, coloration may be darker.

Surface sculpture: Submatt dorsally. As in E. pseudopusilla  sp. n. but microreticulation more weakly impressed, dorsal surface shinier.

Structures: Pronotum with lateral bead. Its lateral sides with distinct longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 slightly truncate.

Male: Antennae simple (Fig. 12). Protarsomere 4 with medium-sized, thick, curved anterolateral hook-like seta. Protarsomere 5 long and narrow, with anterior band of 30 and posterior row of 7 relatively long setae (Fig. 36D). Median lobe in lateral view slightly curved, with slightly thickened, straight apex; in ventral view, subparallel, very slightly narrowed distally, with broad, truncate apex. Paramere slightly concave on dorsal side, with long, dense subdistal setae, median and proximal ones finer and sparser (Fig. 36 A–C). Abdominal ventrite 6 with 4-6 lateral striae on each side.

Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.


Exocelina cyclops  sp. n. is similar to E. sumokedi  sp. n., E. adelbertensis  sp. n., and E. bewani  sp. n., but has coarser dorsal punctation. In this character, the species is more similar to E. pusilla  sp. n. and E. pseudopusilla  sp. n. but it differs from them in much broader and differently shaped median lobe (not distinctly narrowed distally, with apex thicker in lateral and ventral views) and from the latter, also in smaller size and slightly shinier dorsal surface.


Papua: Jayapura Regency. The species is known only from the Cyclops Mountains (Fig. 50).


The species is named after the Cyclops Mountains. The name is a noun in the nominative singular standing in apposition.