Cacostola howdenae Wappes and Santos-Silva,

Wappes, James E. & Santos-Silva, Antonio, 2019, New species and taxonomic notes for Cacostola Fairmaire and Germain, 1859 (Coleoptera: Cerambycidae: Lamiinae: Onciderini), Insecta Mundi 741 (741), pp. 1-20: 7-8

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Cacostola howdenae Wappes and Santos-Silva

new species

Cacostola howdenae Wappes and Santos-Silva  , new species

( Fig. 20–23View Figures 20–27)

Description. Male. Integument mostly dark brown; anteclypeus and labrum reddish-brown; apex of last palpomeres yellowish-brown; basal half of antennomere IV, and basal third of antennomere V reddish-brown; basal 2/3 of antennomere VI yellowish-brown; antennomeres VIII–X yellowish-brown, except narrow dark brown apex; antennomere XI yellowish-brown; elytra with wide dark reddish-brown band dorsally, oblique from humerus to near middle, then narrowed, following parallel to suture and gradually disappearing toward distal quarter; trochanteres reddish-brown; basal third of tibiae dark reddish-brown (less so depending on light intensity); tarsomere V dark reddish-brown, apex dark brown.

Head. Frons finely, moderately abundantly punctate; with yellowish-brown pubescence not obscuring integument, slightly denser laterally and centrally along superior area of median groove, with short white setae interspersed. Vertex with sculpturing as on frons; with dense yellowish-brown pubescence between antennal tubercles, close to upper eye lobes and centrally, remaining surface brown, with short white setae interspersed. Area behind upper eye lobes with sculpturing as on vertex; with dense yellowish-brown pubescence close to superior area of eye, with short white setae interspersed, brown, nearly obscuring integument on remaining surface. Area behind lower eye lobes with sculpturing as on vertex; with dense yellowish-brown pubescence, with short white setae interspersed on superior area close to eye and posterior area close to prothorax; remaining surface with sparse yellowish-brown pubescence and short white setae interspersed. Genae 0.7 times length of lower eye lobe; with sculpturing as on frons, distal area smooth; with sparse yellowish-brown pubescence and short white setae interspersed. Antennal tubercles with inner projection distinctly elevated, narrow; together, in frontal view, U-shaped; sculpturing as on frons basally, minutely punctate on remaining surface; with yellowishbrown pubescence not obscuring integument and short white setae interspersed. Wide central area of postclypeus with sculpturing and pubescence as on frons, smooth, glabrous laterally. Labrum with sparse yellowish-white pubescence close to anteclypeus, slightly denser, with white setae interspersed on anterior area. Mandibles with moderately sparse yellowish-brown pubescence on basal 2/3 of outer surface. Distance between upper eye lobes 0.34 times length of scape; in frontal view, distance between lower eye lobes 0.76 times length of scape. Antennae 2.1 times elytral length, reaching elytral apex about middle of antennomere VII. Scape with sparse yellowish-brown pubescence, slightly denser basally and short white setae interspersed. Pedicel with sparse yellowish-brown and yellowish-white pubescence basally, remaining surface brown. Antennomere III with yellowish-white pubescence on basal third, remaining surface brown; antennomeres IV–VI, with yellowish-white pubescence on light area, brown on dark area; antennomere VII with sparse yellowish-white pubescent maculae; antennomeres VIII–XI with yellowish-white pubescence; pedicel and antennomeres III–VIII with long, erect, thick dark setae ventrally. Antennal formula (ratio) based on length of antennomere III: scape = 0.72; pedicel = 0.15; IV = 1.25; V = 1.12; VI = 1.00; VII = 0.87; VIII = 0.80; IX = 0.67; X = 0.65; XI = 0.80.

Thorax. Prothorax wider than long; sides with distinct rounded tubercle from about middle to posterior constriction. Pronotum coarsely, sparsely punctate; transversely rugose centrally, from about anterior constriction to middle; with white pubescent band centrally on posterior quarter, surrounded by yellowish-brown pubescence; white pubescent on center of posterior margin; with moderately dense yellowish-brown and white pubescence mixed laterally; sparse yellowish-brown pubescence anteriorly and short white setae interspersed; remaining surface with brown pubescence not obscuring integument, with short white setae interspersed. Sides of prothorax with sculpturing as on pronotum; with yellowishbrown pubescence, denser toward prosternum and with short white setae interspersed. Prosternum with dense yellowish-brown pubescence laterally, with white setae interspersed, distinctly sparser centrally. Prosternal process with yellowish-brown pubescence, with white pubescence interspersed, slightly denser than on central area of prosternum; narrowest area of prosternal process about 0.2 times width of procoxal cavity. Mesanepisternum, mesepimeron, sides of mesoventrite, and base of metanepisternum with dense yellowish-brown pubescence; central area of mesoventrite with yellowish-brown pubescence, with white pubescence interspersed; remaining surface of metanepisternum and metaventrite with brownish pubescence not obscuring integument (distinctly sparser on posterocentral area of metaventrite), except sparse yellowish-brown pubescence on metaventrite close to meso- and metacoxae. Scutellum with sparse yellowish-brown pubescence. Elytra. Humeri slightly projected; sides slightly convergent from humeri to middle, parallel-sided from middle to distal sixth, then gradually narrowed toward rounded apex; with two moderately elevated carina from base to near apex, fused posteriorly; sides of posterior sixth depressed in relation to the dorsal surface; with white pubescent band dorsally, from base to posterior sixth, fused with another white pubescent band, also dorsally, from after middle to near apex; sides of dorsal surface with yellowish-brown and white pubescence mixed on anterior third; area between white pubescent band and suture with yellowish-brown pubescence not obscuring integument, except white pubescent band close to suture on about central third; inclined area with sparse yellowish-brown pubescence basally and about middle; remaining dorsal surface with brown pubescence not obscuring integument, sparse yellowish-brown pubescent laterally from middle. Legs. With yellowish-brown pubescence not obscuring integument; meso- and metatibiae tumid, especially the latter.

Abdomen. Ventrites with yellowish-white pubescence, denser laterally, yellower on ventrite V. Apex of ventrite V slightly concave.

Dimensions (mm). Total length, 10.00; prothoracic length, 1.35; anterior prothoracic width, 1.55; posterior prothoracic width, 1.60; maximum prothoracic width, 1.75; humeral width, 2.50; elytral length, 7.50.

Type material. Holotype male from GUATEMALA, Guatemala: 14.5 km SE Guatemala City, Puerta Parada , 1790 m, 13.VI.1991, A. Howden col. ( FSCA, formerly ACMT). 

Remarks. Cacostola howdenae  sp. nov. is similar to C. mexicana (Breuning, 1943)  . As we do not know the sex of the holotype of C. mexicana  , it is not possible to provide a detailed comparison, but the new species at least differs as follows: elytra more distinctly narrowed toward apex, which is also narrower, and not widened after middle; distance between upper eye lobes smaller, distinctly shorter than 1/3 of the basal width of the prothorax (equal to 1/3 of basal width of the prothorax in C. mexicana  ).

Etymology. Named for Anne Howden (now deceased), a wonderful lady and the collector of the holotype.


Florida State Collection of Arthropods, The Museum of Entomology