Gerris (Gerris) costae fieberi Stichel, 1938

Fent, Meral, Kment, Petr, Çamur-Elipek, Belgin & Kirgiz, Timur, 2011, Annotated catalogue of Enicocephalomorpha, Dipsocoromorpha, Nepomorpha, Gerromorpha, and Leptopodomorpha (Hemiptera: Heteroptera) of Turkey, with new records 2856, Zootaxa 2856 (1), pp. 1-84 : 44

Gerris (Gerris) costae fieberi Stichel, 1938

Material examined. EUROPEAN TURKEY: Kırklareli province: Dereköy—Çağlayan (loc. 3), 1 ♂; Kofçaz 10 km under brook (loc. 4), 2 ♀♀ 1 larva, M. Fent det. ( TUET) . Tekirdağ Province: Malkara-Izgar village (loc. 36), fountain, 24.iii.2010, 1 ♀, M. Fent lgt. & det. ( TUET) . ASIAN TURKEY: Aksaray province: Güzelyurt env. (N 38°17' E 34°23'), road from Çiftlik to Güzelyurt , 28.iv.2004, 1 ♀ (ma), P. Kment lgt. & det. ( NMPC) . Ardahan province: 12 km E ŞavŞat, Çam Geçidi [pass] (N 41°12.0' E 42°30.6'), 2430–2485 m a.s.l., steppes and pastures on N slope, ruins of stony shelters near the stream, small tanks, 2.–3.vii.2004, 1 ♂ (ma), J. Hájek & J. Růžička lgt., P. Kment det. ( NMPC) . Artvin province: Barhal [= Altıparmak] (N 40°58'07.2" E 41°24´06.5"), pool under the village, 13.v.2005, 2 ♂♂ 8 ♀♀ (ma), P. Kment lgt. & det. ( NMPC) ; 8 km E ŞavŞat, 1930 m a.s.l., Karagöl Sahara Nat. Park (N 41°13.5' E 42°27.1'), 4.–5.vii.2004, 1 ♂ 1 ♀ (ma), J. Hájek & J. Růžička lgt., P. Kment det. ( NMPC) . Çorum province: Boğazkale env., Yazılıkaya (N 40°01'32.8" E 34°37'54.1"), 1160 m a.s.l., small pool by parking place, 18.v.2005, 2 ♂♂ (ma), P. Kment lgt. & det. ( NMPC) ; Seyfe Çayı [stream], 12.viii.1987, 1 ♂, H. Güher lgt., M. Fent det. ( TUET) . Erzincan province: Çağlayan (N 39°36' E 39°42'), valley above waterfall, brook, 23.vi.2002, 5 ♂♂ 1 ♀♀ (ma), P. Kment lgt. & det. ( NMPC) . Mersin province: Arslanköy env. (N 36°59'26.5" E 34°16'44.5"), 1379 m a.s.l., valley of the brook above the village, 6.v.2007, 1 ♂ (ma), P. Kment lgt. & det. ( NMPC) . Kastamonu province: Kastamonu env., water trough, 30.viii.1970, 2 ♂♂ (ma), D. Goddard lgt., P. Kment det. ( NMPC) . Sivas province: Zara env., Demiryurt (N 39°50'35.1" E 37°36'47.5"), river above the village, 26.vi.2002, 2 ♀♀ (ma), 16.v.2005, 3 ♂♂ 2 ♀♀ (ma), P. Kment lgt. & det. ( NMPC) . Tunceli province: Tunceli (N 39°06' E 39°33'), Munzur Nehri [river] valley, oxbow lake near bridge, 22.vi.2002, 1 ♂ (ma), P. Kment lgt. & det. ( NMPC) .

European Turkey. Josifov (1986a), Andersen (1995), Önder et al. (2006, as G. costae ), Aukema (2009); this paper. First exact records from Turkish Thrace.

Asian Turkey. Horváth (1883, as G. costae ), Oshanin (1908, as Gerris (Gerris) costae ), Oshanin (1912, as G. costae ), Kiritshenko (1918, as G. lateralis , misidentifications), Poisson (1925, as G.costae ), Gadeau de Kerville (1939, as G. Costae ), Hoberlandt (1952a, as G. costae ; revised!), Andersen (1995), Kıyak et al. (2004, 2008, all as G. costae ), Damgaard (2006, as G. costae ), Önder et al. (2006, as G. costae costae ), Salur & Mesci (2009, as G. costae ).

Turkey (not distinguished). Stichel (1955, as G. costai fieberi ), Kanyukova (1982, as G. costae costae ), Nieser & Moubayed (1985), Kıyak & Özsaraç (2001), Kanyukova (2006).

General distribution. Gerris costae fieberi is distributed in Italy, Balkan Peninsula, Ukraine, Central and South European Territory of Russia, European part of the Kazakhstan, Egypt, Anatolia, Transcaucasia, Israel, Lebanon, Syria, Iraq, Iran, Afghanistan, Turkmenistan, and Uzbekistan ( Andersen 1995, Protić 1998, Saleh Ahmed & Gadalla 2005). Gerris c. costae (Herrich-Schaeffer, 1850) is limited to the Alps and G. c. poissoni Wagner & Zimmermann, 1955 to western Europe ( Andersen 1995). Gheit (1995) reported G. costae from Morocco without indication of the subspecies. Klingenberg (1992) confirmed the validity of the subspecies by multivariate morphometric study. However, Damgaard (2006), based on molecular data, downgraded Gerris sahlbergi Distant, 1879 (Central Asia, Siberia, Mongolia, China, northern India) to a subspecies of G. costae , and further questioned the validity of the included subspecies based on the lack of geographical substructure in the mitochondrial DNA examined.

Identification. For identification of the species see Kanyukova (1982, 2006), Andersen (1994), Linnavuori (1998), Linnavuori & Hosseini (2000). Diagnoses of the subspecies are given by Wagner & Zimmermann (1955).