Clistopyga crassicaudata Sääksjärvi, Bordera & Palacio

Palacio, Edgard, Bordera, Santiago, Sääksjärvi, Ilari E. & Díaz, Francisco, 2018, The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part II: the C. isayae species group, with the description of seven new species, Zootaxa 4442 (1), pp. 101-121: 103-106

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Clistopyga crassicaudata Sääksjärvi, Bordera & Palacio

sp. nov.

Clistopyga crassicaudata Sääksjärvi, Bordera & Palacio  sp. nov.

( Figs 1B View Figure , 2A–F View Figure , 9A View Figure )

Diagnosis. Clistopyga crassicaudata  sp. nov. can be distinguished from all other species of the C. isayae  species group by the combination of the following characters: upper valve of ovipositor distinctly thickened apically ( Fig. 1B View Figure ); ocular orbits widely interrupted laterally and posteriorly ( Figs 2A, B View Figure ); subalar prominence sparsely punctuate, with about 15˗17 setiferous punctures; occipital carina present only ventrally as a short vestige; clypeus 2.0 × as broad as medially long; tergite I 1.3 × as long as posteriorly broad ( Fig. 9A View Figure ); hind wing with abscissa of Cu 1almost straight ( Fig. 2F View Figure ).

Description. Female: Body length 9.8 mm. Fore wing length 6.5 mm.

Head ( Figs 2A–B View Figure ). In dorsal view, moderately narrowed behind eyes. Gena smooth and shiny with uniformly sparse setiferous punctures, in dorsal view 0.26 × as long as eye, in frontal view almost straight and moderately constricted below eyes. Frons smooth and shiny. Vertex smooth and shiny, with isolated setiferous punctures. Posterior ocellus separated from eye 0.81 × its maximum diameter ( Fig. 2B View Figure ). Distance between hind ocelli 1.0 × maximum diameter of posterior ocellus. Occipital carina present only ventrally as a short vestige. Face with fine and dense setiferous punctures, distance between punctures more than twice the diameter of punctures. Clypeal suture strongly curved. Clypeus 2.0 × as broad as medially long, distinctly convex in dorsal half, flat in ventral half, with apical margin bilobate ( Fig. 2A View Figure ). Malar space 0.8 × as long as basal mandibular width, with a granulate stripe between eye and mandible. Antenna with 25 flagellomeres, first flagellomere about 8.0 × as long as wide.

Mesosoma  ( Figs 1B View Figure , 2D View Figure ). Pronotum shiny, mostly smooth, with fine and scattered setiferous punctures in upper posterior part. Mesoscutum entirely smooth and shiny, with fine and sparse setiferous punctures. Notauli deep, reaching about 0.4 of length of mesoscutum. Mesopleuron shiny, with moderately sparse setiferous punctures, except in posterior part under speculum. Subalar prominence sparsely punctate, with about 15˗17 setiferous punctures. Epicnemial carina weak, its dorsal end straight, ending slightly below level of centre of pronotum. Metapleuron shiny, glabrous, except for some sparse setiferous punctures at posterior end, 2.5 × as long as deep. Propodeum smooth and shiny, with sparse and fine setiferous punctures laterally; in dorsal view 1.2 × as long as medially wide. Propodeal spiracle contiguous with groove separating propodeum and metapleuron, groove not interrupted by spiracle ( Fig. 1B View Figure ). Hind leg with femur about 4.5 × as long as deep, about 0.90 × as long as tibia.

Wings ( Figs 2E–F View Figure ). Fore wing with vein cu-a slightly distal to Rs & M. Vein 2 rs-m about 0.4 × as long as abscissa of M between 2 rs-m and 2 m-cu. Abscissa of Cu 1 between 1 m-cu and Cu 1 a 1.7 × as long as Cu 1 b ( Fig. 2E View Figure ). Hind wing with vein cu-a about 0.24 × as long as abscissa of Cu 1 between M and cu-a ( Fig. 2F View Figure ). Vein cu-a reclivous, abscissa of Cu 1 vertical and almost straight. Vein Cu 1 well pigmented.

Metasoma ( Figs 1B View Figure , 2C View Figure , 9A View Figure ). Tergite I 1.3 × as long as posteriorly broad ( Fig 9A View Figure ), smooth and shiny, with fine and relatively dense setiferous punctures laterally; spiracle near its basal 0.4; lateromedian longitudinal carinae strong, reaching about 0.2 of length of tergite; lateral longitudinal carinae weak, reaching about 0.4 of length of tergite. Sternite I extending back about 0.5 of length of tergite. Tergite II 1.20 × as long as posteriorly broad ( Fig. 2C View Figure ), central region shiny, with very fine and moderately sparse setiferous punctures; rest of tergites shiny, progressively more densely and strongly punctate. Ovipositor very stout, upper valve distinctly thickened apically ( Fig. 1B View Figure ), weakly upcurved at distal 0.2, 3.0 × as long as hind tibia. Ovipositor sheath about 2.4 × as long as hind tibia, length of setae on average about 1.5 × sheath basal width.

Colouration. Head blackish brown, with clypeus, frontal orbits, face orbits widely, mouth parts except apex of mandibles white ( Figs 1B View Figure , 2A, 2B View Figure ); antenna brown, basal flagellomeres, pedicel and scape pallid on the outer side. Mesosoma  mostly orange ( Fig. 1B View Figure ), scutellum posteriorly and metanotum, white, propleuron, axillae of mesonotum and metanotum, propodeal furrow, propodeum except laterally stripes, blackish brown. Metasoma brown, tergites I˗IV with white blotches near the posterior corners ( Fig. 2C View Figure ), tergites II˗VI with posterior corners marked with black ( Figs 1B View Figure , 2C View Figure ); ovipositor sheath basally brown and apically pallid. Fore and mid legs predominantly white, coxae, trochanteral segments and femora dorsally brown, tibiae ventrally brown, tarsi brownish; hind leg predominantly brown, coxa dorsally and ventrally, trochanter distally, femur ventrally and laterally, tibia laterally on the middle, white; hind tarsal segments brown but basally white. Wings hyaline, pterostigma brown.

Male: unknown.

Etymology. The specific name refers to the distinctly thickened ovipositor apex.

Type material. Holotype 1 ♀: Perú, Dept, of Loreto, Río Sucusari, Río Napo , Explornapo Camp , 0 3o 15´S, 72o55´W, 100m, hand collecting/sweeping, 3 Jun 1992, coll. T. L. Erwin ( ZMUTAbout ZMUT). 

Distribution: Perú.


University of Tokyo, Department of Zoology