Orphnabaccha Hull, 1949

Mengual, Ximo, Miranda, Gil Felipe G. & Thompson, F. Christian, 2018, Unraveling Ocyptamus and the Baccha legacy (Diptera: Syrphidae): redefinition of groups and new species descriptions, Zootaxa 4461 (1), pp. 1-44 : 13

Genus Orphnabaccha Hull, 1949 stat. rev.

Figs 8 , 55–60 , 104, 105 .

Orphnabaccha Hull, 1949a: 93 , as a genus. Type species: Baccha coerulea Williston, 1891 (original designation).

Mercurymyia Fluke, 1950a: 140, as a subgenus of Stenosyrphus Matsumura & Adachi, 1917 (jun. syn. of Melangyna Verrall ). Type species: Epistrophe caldus Walker, 1852 (original designation). Synonymy by Thompson et al. (1976: 11).

Ocyptamus caldus species group. Thompson (1981).

Ocyptamus (Orphnabaccha) . Mengual et al. (2012).

Ocyptamus amplus species group. Miranda et al. (2016).

Diagnosis. Frons wide. Ocellar triangle distanced three ocellus-width from the eye margin. Dorsal lobe of calypter with long pile. Metasternum usually pilose. Usually with parallel-sided elongated to slightly oval abdomen, some species slightly petiolate.

Comments. Flies from this group usually have broader abdomens, and some species are very Syrphus -like. Others are darker, with shiny flattened pile on the abdomen. Not all Orphnabaccha species have the distinctive pilose metasternum, but they can be further distinguished from the other groups presented here by a usually broader face, female ocellar triangle distinctly distanced from the eye margin and dorsal lobe of the calypter with long pile. Currently, there are three species groups within the genus, each with distinct types of genitalia ( Vockeroth 1969): 1) O. ampla species group ( Syrphus -like flies with bifurcate postgonites); 2) O. calda species group (species with arched yellow fascia on the abdomen, postgonite tooth-like, and distiphallus with large spines); and 3) O. coerulea species group (black abdomen with shiny flattened pile and lamellate postgonites).

Species list (31 species): O. aequilineata ( Hull, 1945) , O. agilis ( Bigot, 1884b) , O. ampla ( Fluke, 1942) , O. calda ( Walker, 1852) , O. coerulea ( Williston, 1891) , O. decipiens ( Williston, 1891) , O. delimbata ( Enderlein, 1938) , O. diversa ( Williston, 1891) , O. dolorosa ( Hull, 1950) , O. elegans ( Giglio-Tos, 1892) , O. erratica ( Williston, 1888) , O. flavigaster ( Hull, 1944d) , O. fraterna ( Bigot, 1884a) , O. fuscicosta ( Lynch Arribálzaga, 1891) , O. golbachi ( Fluke, 1950b) , O. jactator ( Loew, 1861) , O. lanei ( Fluke, 1950b) , O. laticauda ( Curran, 1941) , O. lativentris ( Curran, 1941) , O. lauta ( Giglio-Tos, 1892) , O. limbus ( Enderlein, 1938) , O. nodosa ( Hull, 1930) , O. opaca ( Fluke, 1950b) , O. priscilla ( Hull, 1943b) , O. pteronis ( Fluke, 1942) , O. superba ( Thompson, 1981) , O. tiarella ( Hull, 1944b) , O. trabis ( Fluke, 1942) , O. tribinicincta ( Enderlein, 1938) , O. virga ( Fluke, 1942) , O. volcana ( Fluke, 1942) .

Figures

FIGURES 1–10. 1: Mimocalla erebus (Hull) [female, CNC_Diptera209354], metafemur, lateral view. 2: Salpingogaster sp. [male, INPA-DIP0000125], metafemur, lateral view. Note: Arrow indicates ventral spinose setae. 3: Salpingogaster nigra Schiner [male, ZFMK-DIP-00020899], postmetacoxal bridge with tergum 1 and sternum 1, posterior view. Note: Arrow indicates tergum 1. Red lines indicate the complete postmetacoxal bridge (=metepimera continuous behind coxae). 4: Eosalpingogaster umbra Mengual & Thompson [female, USNMENT 00036793], postmetacoxal bridge with tergum 1 and sternum 1, posterior view. Note: Arrow indicates tergum 1. Red lines indicate the complete postmetacoxal bridge (=metathoracic epimera continuous behind metacoxae); less sclerotized medial line visible. 5: Salpingogaster nigra [male, DEBU01088906], wing. 6: Eosalpingogaster cochenillivora (Guérin-Méneville) [male, MPEGDIP12170472], wing. 7: Pelecinobaccha alia Miranda [male, USNMENT 01288155], occiput and pleuron, lateral view. Note: Arrow indicates pile on anterior anepisternum. 8: Orphnabaccha cf. ampla Fluke [male, CNC_Diptera210129], pleuron, lateral view. Note: Arrow indicates pile on anterior anepisternum. 9: Salpingogaster nigra [male, ZFMK-DIP-00020899], scutellum and tergum 1, dorsal view. 10: Eosalpingogaster umbra [female, USNMENT 00036793], scutellum and tergum 1, dorsal view.

FIGURES 55–61. 55: Orphnabaccha lanei (Fluke) [male paratype, MZUSP], habitus, dorsal view. 56: Orphnabaccha coerulea (Williston) [male, USNM], habitus, dorsal view. 57: Orphnabaccha coerulea [male, USNM], head, frontal view. 58– 60: male genitalia: a: lateral view; b: hypandrium, postgonite and phallus, ventral view; c: postgonite, lateral view, outline only; d: distal portion of phallus, anteroventral view. 58: Orphnabaccha calda (Walker) (from Vockeroth 1969). 59: Orphnabaccha coerulea, male genitalia (from Vockeroth 1969). 60: Orphnabaccha ampla (Fluke), male genitalia (from Vockeroth 1969). 61: Pipunculosyrphus globiceps Hull [female, ZFMK], habitus, dorsal view. Scale = 1 mm.

FIGURES 99–105. 99: Pelecinobaccha summa (Fluke) [female, CNC], head and thorax, dorsolateral view. 100: Pelecinobaccha adspersa, female, by Gil F. G. Miranda. 101: Pelecinobaccha impostor Miranda [female, ZFMK-DIP- 00018039], 6 th abdominal segment female, lateral view. Note: Arrow indicates the lateral weakness line. 102: Pelecinobaccha adspersa [female, DEBU00186967], 5 th and 6 th abdominal segment female, lateral view. Note: Arrow indicates the 6 th abdominal segment.103: Pelecinobaccha pilipes (Schiner) [female, USNMENT 00257696], metaleg, lateral view, photo previously published in Miranda et al. (2014). 104: Orphnabaccha cf. lativentris (Curran) [male, CNC, JSS25215], habitus, lateral view. Scale = 1 mm. 105: Orphnabaccha coerulea [male, USNM], habitus, lateral view.