Ocyptamus maximus Thompson

Mengual, Ximo, Miranda, Gil Felipe G. & Thompson, F. Christian, 2018, Unraveling Ocyptamus and the Baccha legacy (Diptera: Syrphidae): redefinition of groups and new species descriptions, Zootaxa 4461 (1), pp. 1-44: 15-16

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Ocyptamus maximus Thompson

sp. nov.

Ocyptamus maximus Thompson  sp. nov.

Figs 11 View Figure , 28 View Figure , 32–37 View Figure , 113.

Ocyptamus  CR-45 Thompson in litt.

Ocyptamus maximus Thompson  in litt. Arcaya et al. (2013).

Differential diagnosis. Although described as Ocyptamus  , this species is incertae sedis regarding its generic affinities (see Comments below). Very large fly, almost completely dark with tawny wings and hyaline costal area. Abdomen petiolate, dark, tergum 2 with two anterolateral yellow/translucent maculae and two medial, divergent orange fasciate markings.

Description. MALE: Head ( Figs 28 View Figure , 32, 34 View Figure ). Face brownish black except for very indistinct orange vitta running from antennal base to level of tubercle, sparsely grayish microtrichose, slightly denser laterally, black pilose; gena orange, shiny and bare on anterior 2/3, brown, dull and black pilose posteriorly; mouthparts orange; occiput black, densely grayish-white microtrichose, white pilose on ventral half, black pilose dorsally, with pile in 3–4 rows at medial notch, and a single row on dorsal 1/5; frontal triangle black, shiny and bare on anterior 1/2, brownish-black microtrichose and black pilose posteriorly; vertical triangle equilateral, black, black microtrichose and pilose; antenna brown, black pilose; ratio 1:1:2, with basoflagellomere about twice as long as broad; arista bare, about 1/3 shorter than antenna. Thorax ( Figs 32, 33 View Figure ). Dark brown except postpronotum and scutellum lighter brown (almost tan); scutum brown microtrichose with lighter brown medial and submedial microtrichose vittae, black pilose, without any distinct anterior pile; scutellum light brown microtrichose, black pilose, without ventral pile fringe; pleuron dull, densely gray microtrichose on katepisternum and posterior anepisternum, black pilose; plumula short, light brown; calypter brownish black, with margin and fringe black, pile long on both calypters; halter and capitulum pale; metasternum bare; metathoracic episternum black pilose. Legs: brownish black except femoral-tibial joints lighter orange brown, black pilose. Wing: microtrichose, light brown except darker on basal cells, cell r1 and r2+3; alula normal, broad, about as broad as cell cup. Abdomen ( Figs 32, 33, 35 View Figure ). Petiolate, with 2nd segment about half as wide medially as apically; tergum 1 brownish black, dull, black pilose; tergum 2 brownish black except for translucent yellow divergent yellow vitta on basal 2/3 and with similar but smaller brownish orange divergent vitta posterior to the yellow and narrowly separated by black color, black pilose; terga 3–5 black, black pilose; sterna 1 and 3–5 black, black pilose; sternum 2 translucent yellow on basal 2/3, black apically, black pilose; 8th segment with a small protuberance. Genitalia: small, cercus orange, as in fig. 113.

FEMALE ( Figs 36, 37 View Figure ). Similar to male except normal sexual dimorphism, frons black, shiny on anterior 1/3, black microtrichose with lighter gray microtrichose medial vitta, black pilose; tergum 6 short, about 1/3 as long as tergum 5, black, black pilose; ovipositor orange.

Length (n=1). Body, 22.4 mm; wing, 16.8 mm.

Geographical distribution. Colombia, Costa Rica and Venezuela.

Type-locality. COSTA RICA: Puntarenas Prov., Parque Nacional La Amistad, Zona Protectora Las Tablas, Las Alturas de Cotón , Estación Biológica Las Alturas , 1569 m., 8°56'43'' N 82°50'00'' W.GoogleMaps 

Material examined. Holotype, male, deposited in the Instituto Nacional de Biodiversidad ( INBioAbout INBio), Santo Domingo de Heredia, Costa Rica, and labelled: “ COSTA RICA. Prov   . Puntarenas. Z. P. // Las Tablas, Estacion Biológica Cotón. // 1569m. 22 JUL 1999. M. Alrao. // Manual L _S_320400_593800 #57455” “ INB003309582View Materials // INBIOCRI COSTA RICA ” [barcode] “ Holotype // Ocyptamus maximus  // Thompson 2013 ” [light red, handwritten] ( Figs 32, 33 View Figure )  . Paratypes: COLOMBIA: Sierra Nevada , San Lorenzo, J. Ujhelyi (1♀ FSCAAbout FSCA)  . COSTA RICA: Puntarenas Prov., Finca Cafrosa, Embalse, 800 m north de Tigra , L_S_317800_ 596200, 1280 m, 15.viii.1997, A. Picado, #47694 (1♀ USNMAbout USNM, INBIOCRI002559410 INBIO)  ; Puntarenas Prov., Sabalito, Camino a la Neblina , L_S_317800_ 597500, 1340 m., 22.viii.2000, M. Alfaro, #58936 (1♂ USNMAbout USNM, INB0003306083View Materials INBIOCRI) ( Figs 11 View Figure , 28 View Figure , 34, 35 View Figure )  . VENEZUELA: Lara, P.N. Terepaima, 1200 m., 6.viii.1976, J.M. Osorio (1♀ MJMOAbout MJMO, CE-UCLA /DS-1235) ( Figs 36, 37 View Figure ). 

Etymology. The specific epithet is derived from the Latin maximus  , which means ‘greatest’ ( Brown 1956: 516), as this is the largest known species of Ocyptamus  and of the subfamily Syrphinae  . Species epithet is an adjective.

Comments. Ocyptamus maximus  is readily recognizable as this is the largest known species of the subfamily Syrphinae  . Only some species of the Eristaline genus Milesia  are larger.

O. maximus  seems to be more morphologically closely related to sargoides Macquart  than to any other taxon listed here based on a similar pattern of translucent maculae on the tergum 2 and sternum 2, the long black pile on the dorsal lobe of the calypter, and male genitalia (similar overall shape of phallus, postgonite and surstylus). Although sargoides Macquart  is currently considered incertae sedis (see below), this new species is placed in the genus Ocyptamus  at the present time since a species name must contain a generic name and a specific epithet [Article 5.1, International Code of Zoological Nomenclature (1999)].


National Biodiversity Institute, Costa Rica


Florida State Collection of Arthropods, The Museum of Entomology


Smithsonian Institution, National Museum of Natural History


Universidad Centro Occidental, Decanato de Agronomia