Belisana cucphuong, Zhu & Li, 2021

Zhu, Wenhui & Li, Shuqiang, 2021, Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia, Zootaxa 4963 (1), pp. 115-137 : 119-122

publication ID	https://doi.org/ 10.11646/zootaxa.4963.1.5
publication LSID	lsid:zoobank.org:pub:6B4B8F71-7B08-4027-A76B-7EE8C987C9BD
DOI	https://doi.org/10.5281/zenodo.4696998
persistent identifier	https://treatment.plazi.org/id/4E3C87F3-FFC5-FF96-C6EC-FCE8FC3AFF7A
treatment provided by	Plazi
scientific name	Belisana cucphuong
status	sp. nov.

Treatment

Belisana cucphuong View in CoL sp. nov.

Figs 3–4 View FIGURE 3 View FIGURE 4 , 13C–D View FIGURE 13 , 14C–D View FIGURE 14 , 15B View FIGURE 15

Type material. Holotype: Male ( IZCAS Ar41904), Cuc Phuong National Park (20°20’58.32”N, 105°35’36.00”E, elevation 437 m), Ninh Binh, Vietnam, 20 August 2015, D.S. Pham leg. GoogleMaps Paratypes: 2 females ( IZCAS Ar41905– 41906), same data as holotype GoogleMaps .

Etymology. The specific name refers to the type locality and is a noun in apposition.

Diagnosis. This species resembles B. vietnamensis Yao, Pham & Li, 2015 (see Yao et al. 2015: 10, figs 29–31) with the elongated bulbal apophyses ( Fig. 4D View FIGURE 4 ) and the male chelicerae ( Figs. 4C View FIGURE 4 , 15B View FIGURE 15 ), but males can be distinguished by the long distal spine, 7 times as long as wide (arrow 2 in Figs 3C View FIGURE 3 , 13C View FIGURE 13 , 4 View FIGURE 4 times in B. vietnamensis ), on the procursus, and by the presence of a long ventral apophysis on the palpal trochanter (arrow 1 in Fig. 3B View FIGURE 3 ) and distinct ventral apophysis on palpal femur (arrow 3 in Fig. 3B View FIGURE 3 ). Females can be distinguished by the vulval pore plates (arrow pp in Figs 4B View FIGURE 4 , 14D View FIGURE 14 ) being longer than wide (vulval pore plates approximately as wide as long in B. vietnamensis ).

Description. Male (holotype, IZCAS Ar41904): Total length 2.22 (2.35 with clypeus), carapace 0.94 long, 0.78 wide, opisthosoma 1.41 long, 0.78 wide. Leg I: – (5.70 + 0.38 + 5.77 + – + –), leg II: 13.00 (3.97 + 0.32 + 3.33 + 4.10 + 1.28), leg III: 8.28 (2.50 + 0.26 + 1.99 + 2.82 + 0.71), leg IV: 14.15 (6.03 + 0.30 + 2.88 + 3.85 + 1.09); tibia I L/d: 38. Distance PME-PME 0.09, diameter PME 0.09, distance PME-ALE 0.09, AME absent. Habitus as in Figs 4E–F View FIGURE 4 . Carapace pale, with brown lateral margins. Ocular area not elevated. Clypeus unmodified. Chelicerae ( Figs. 4C View FIGURE 4 , 15B View FIGURE 15 ) with pair of small proximo-lateral apophyses and pair of short distal apophyses, each with sclerotized and pointed tip (distance between tips of distal apophyses: 0.04). Sternum pale, approximately as wide as long (0.50). Thoracic furrow absent. Legs pale. Opisthosoma pale, with large, dark spots. Palps as in Figs 3A–B View FIGURE 3 ; coxa unmodified; trochanter with long ventral apophysis (arrow 1 in Fig. 3B View FIGURE 3 ); femur with small proximo-dorsal apophysis (arrow 2 in Fig. 3B View FIGURE 3 ) and distinct ventral apophysis (arrow 3 in Fig. 3B View FIGURE 3 ); procursus ( Figs 3A–D View FIGURE 3 , 13C–D View FIGURE 13 ) simple proximally but complex distally, with large subdisto-prolateral sclerite (arrow 1 in Figs 3C View FIGURE 3 , 13C View FIGURE 13 ), curved distal spine (arrow 2 in Figs 3C View FIGURE 3 , 13C View FIGURE 13 ) and subdisto-retrolateral membranous process (arrow in Figs 3D View FIGURE 3 , 13D View FIGURE 13 ); bulb ( Fig. 4D View FIGURE 4 ) with hooked apophysis and simple embolus. Retrolateral trichobothria of tibia Iat 5% proximally; without spines or curved setae.

Female (IZCAS Ar41906): Similar to male, habitus as in Figs 4G–H View FIGURE 4 . Total length 1.90 (2.00 with clypeus), carapace 0.72 long, 0.66 wide, opisthosoma 1.28 long, 0.66 wide. Tibia I: 4.04; tibia I L/d: 51. Distance PME-PME 0.13, diameter PME 0.06, distance PME-ALE 0.09, AME absent. Sternum slightly wider than long (0.50/0.40). Opisthosoma with indistinct dark spots. External female genitalia ( Figs 4A View FIGURE 4 , 14C View FIGURE 14 ) simple and flat, with pair of frontal “epigynal” pockets on sclerotized plate (arrow ep in Figs 4A–B View FIGURE 4 , 14C–D View FIGURE 14 ), 0.03 apart. Vulva ( Figs 4B View FIGURE 4 , 14D View FIGURE 14 ) with wavy anterior arch (median ridge strongly protruding, arrow in Figs 4B View FIGURE 4 , 14D View FIGURE 14 ) and pair of nearly elliptical pore plates (arrow pp in Figs 4B View FIGURE 4 , 14D View FIGURE 14 ), without serrated sclerites.