Sirindhornia Pinkaew and Muadsub, 2014

Muadsub, Sopita & Pinkaew, Nantasak, 2014, Sirindhornia Pinkaew and Muadsub (Lepidoptera: Tortricidae), a new enarmoniine genus from Thailand, Zootaxa 3869 (1), pp. 53-63 : 54-55

Sirindhornia Pinkaew and Muadsub , n. gen.

Type species: Sirindhornia pulchella Pinkaew and Muadsub , n. sp.

Diagnosis. Sirindhornia is superficially most similar to Anthozela Meyrick but can be distinguished by its upcurved and appressed, long, slender, white labial palpi, by a forewing with white rather than yellow ground color in its basal half, and a venation similar to that of Irianassa Meyrick with CuA 1 from far below the angle of the cell in the forewing and R s and M 1 widely distant at base in the hindwing. The male genitalia are most like those of Anthozela , but those of Sirindhornia have large spiny socii with 2–3 huge apical thorns, long slender sclerotized gnathos arms, and a long slender henion. The large subtriangular blade-shaped signa of the female are typical of many Enarmoniini are unlike the long slender hooks of Anthozela .

Description. Head: Labial palpi long, slender, upcurved, reaching upper margin of eyes, close to frons ( Fig. 1 ); frontal tuft small, with scales on lower frons short, appressed, white; vertex with raised black and white scales; labial palpus with short basal segment with raised scales, also laterally, white except for laterodistal black scales; second segment strongly curved, widest in middle, with long anteriorly projecting transverse scales, white except for black scales basally; third segment long, straight, pointed, cone-shaped with deeply concave anterior face, white, in some species with black scales in ventral groove. Antenna short, 1/3 length of forewing, scape and pedicel black, flagellum with dorsal scales only, black with white scales in some species, sensory setae minute. Ocellus prominent, chaetosema with long sensory setae.

Thorax: Smooth, without raised scale tufts posteriorly, black and white; legs unmodified. Forewing ( Fig. 3 ) broadly triangular, length 3.8–5.2 mm in male (n = 5), 4.5–5.3 mm in female (n = 4); costa evenly curved, rarely more so before apex, male costal fold absent, termen with notch below apex, wing venation ( Fig. 2 ) with discal cell 0.65x length of wing, M-stem well developed, extending between M 2 and M 3, chorda from halfway between R 1 and R 2 to just beyond R 5; all veins present and separate beyond cell, R 1 from basal 0.33 of cell, R 2 from apical 0.25 of cell, R 3 from much closer to R 4 than to R 2, bases of R 3, R 4 and R 5 nearly equidistant, M 1 from slightly closer to R 5 than to M 2, base of M 2 equidistant to M 1 and M 3, M 3 from angle of cell, base of CuA 1 from 0.75 of cell and equidistant to M 3 and CuA 2, CuA 2 from about middle of cell, 1A and 2A separate at base and connate at 0.38 length of anal vein. Forewing pattern with basal 1/3–2/5 white, with irregular and often confluent black spots, distal part with orange to brownish orange ground color, its inner margin preceded by two dark spots and extending obliquely from basal 0.2 of costa to R 2, then angled and roughly straight to dorsum; costa with well developed strigulae as black spots alternating with white along costa, giving rise to strongly oblique lines, an angled silvery one from 1/3 costa separating white and red wing portion, an irregular, nearly straight black line from before middle to near notch on termen, and a shorter, sinuate, silvery line just beyond a black spot on costa; wing apex with two distinct white marks, a triangular one followed by a crescent-shaped one, their margins edged with black and silvery scales; notch below apex often enhanced by a small, triangular, white mark; remainder of orange wing portion with connected, sinuate silvery or blackish lines and few black spots. Fringe blackish from wing apex to CuA 1, in some species interrupted by white at notch between R 5 and M 1, remainder orange brown mixed with blackish, basal line brownish orange. Fringe pale orange grey with conspicuous blackish basal line. Underside dark grey with greyish orange band along termen, in some species with one or two longitudinal, narrowly triangular greyish orange patches. Underside of forewing greyish brown, speckled paler reflecting wing pattern; costa with small white dots and two distal marks as on upper side; with white scales at notch in some species. Hindwing with frenulum comprised of one bristle in male and three bristles in female, R-stem well developed to base, base of M 1 conspicuously distant from R s, M 2 distant from and subparallel with M 3, M 3 and CuA 1 connate to short-stalked, CuA 2 from near apical third, CuP very weak towards margin, 1A+2A and 3A well developed. Hindwing pattern blackish, paler toward base, with orange band along termen, usually with orange patch near center.

Abdomen: Male genitalia ( Fig. 4 ) with tegumen ovate-subtriangular, moderately sclerotized, homology of highly derived dorsal appendages unclear; with flat medial, dorsal process (presumably uncus) and large, curved raised lateral lobes (presumably socii) with dense flattened modified spines on outer surface and with 2–3 huge thorns on process projecting from outer margin, and with curved sclerotized arms along base, their tips with a membranous medial connection (presumably gnathos); vinculum wide, weakly sclerotized band; juxta small, caulis rather short; anellus with long sclerotized spatulate dorsal process, with microtrichia on apex; phallus long, bulbous, without cornuti; valva long and narrow, sacculus from small basal thickening to forming long distally free process, costal hook prominent, pointed, usually below costa, outer surface of valva with long, persistent, modified scales in basal half. Female genitalia ( Fig. 5 ) with papillae anales densely setose; tergum VIII with very sparse scales sockets and setae posteriorly and on lateral triangular extensions; sterigma a large patch of dense spinules, some species with a projecting pyramid-shaped lobe covered with spinules on each side; ostium a membranous to sclerotized funnel beyond posterior margin of sternum VII; without colliculum; ductus bursae long, very narrow in some species, wider near ostium or posterior half, ductus seminalis arising from transition of corpus to ductus bursae; corpus bursae ovate, granulate, with two large unequal signa, triangular blades with elongate hollow base.

Etymology. Sirindhornia is a patronym for Princess Maha Chakri Sirindhorn..

Distribution. Thailand.

Figures

FIGURE 1. Head of Sirindhornia spp. A–B. S. pulchella, n. sp. (holotype male). C–D. S. chaipattana, n. sp. (holotype male). E–F. S. curvicosta, n. sp. (holotype male). G–H. S. bifida, n. sp. (holotype male). I–J. Sirindhornia sp. (female).

FIGURE 2. Wing venation of Sirindhornia pulchella, n. sp. (paratype male, right wings).

FIGURE 3. Adults of Sirindhornia spp. A. S. pulchella, n. sp. (holotype male). B. S. pulchella, n. sp. (paratype female). C. S. chaipattana, n. sp. (holotype male). D. S. chaipattana, n. sp. (paratype female). E. S. curvicosta, n. sp. (holotype male). F. S. bifida, n. sp. (holotype male). G. Sirindhornia sp. (female) (Scale bar = 2 mm).

FIGURE 4. Male genitalia of Sirindhornia spp. A. S. pulchella, n. sp. (holotype). B. S. chaipattana, n. sp. (holotype). C. S. curvicosta, n. sp. (holotype). D. S. bifida, n. sp. (holotype).

FIGURE 5. Female genitalia of Sirindhornia spp. A. S. pulchella, n. sp. (paratype). B. S. chaipattana, n. sp. (paratype). C. Sirindhornia sp.