Lichtwardtia nabozhenkoi Grichanov, 2020

Grichanov, I. Ya., 2020, New species and new records of Lichtwardtia Enderlein, 1912 (Diptera: Dolichopodidae) from Australasian and Oriental regions, Far Eastern Entomologist 399, pp. 1-13 : 6-9

Lichtwardtia nabozhenkoi Grichanov , sp. n.

http://zoobank.org/NomenclaturalActs/ 56E9EC17-94E8-4134-B869-503540443585

Figs 9 – 16

TYPE MATERIAL. Holotype – ♂, Indonesia: Bali Is., Buyan Lake , 8.24º S,

115.14º E, 1220 m, 27.XII 2016, leg. N. Vikhrev [ ZMUM].

wing; 13 – postabdomen; 14 – hypandrium and phallus; 15 – cercus, inner view, surstylus and postgonite; 16 – cercus, outer view.

DESCRIPTION. Male. Head. Frons metallic greenish violet, weakly pollinose;

face entirely white pollinose; one strong vertical, one short postvertical, a pair of strong ocellar setae present; lower postocular setae white; ventral postcranium with

2 long white setae and several light cilia; eyes with short hairs; face glabrous; face almost parallel-sided, narrowest at upper third, slightly widening at clypeus;

clypeus almost straight ventrally, not reaching lower margin of eyes; ratio of its minimal width to height 21/60; antenna mostly yellow-orange; postpedicel black in distal half, rounded, about as long as high (14/15), with short hairs; arista-like stylus middorsal, black, sparsely pubescent, with hairs 2-4 times longer than basal diameter of stylus; length ratio of scape to pedicel to postpedicel to stylus (1st and

2nd segments), 9/8/14/11/44; palpus small, dirty yellow, with with black seta, short light and dark hairs; proboscis brown.

Thorax. Mesonotum metallic dark-blue-violet, weakly brownish pollinose; pleura greenish-black, whitish pollinose; 5 strong dorsocentral setae with several microscopic hairs in front of the 1st pair, 2 rows of short acrostichals; proepisternum with 1 strong black seta above fore coxa and several short hairs; scutellum with 2

strong setae and 2 very short lateral hairs.

Legs mostly yellow; fore coxa yellow, mid coxa brown-black with yellow apex,

hind coxa yellow; tarsi black from tip of basitarsus; femora without long hairs; fore coxa with black hairs and several long apical setae; fore tibia with 1 long and 1 small anterodorsal, 1 long and 1 small dorsal, 1 posterior seta, more than 2 times as long as diameter of tibia, 3 apical setae of about equal length; fore tarsus simple, fore basitarsus with short basoventral seta; mid femur with 1 subapical anterior seta; mid tibia with 3 anterodorsal, 2 posterodorsal, 1 anteroventral and 5 apical setae; hind femur with one anterodorsal prepapical seta; hind tibia simple, with 3-4 anterodorsal,

3-4 posterodorsal, 2-3 ventral, 3 apical setae; hind basitarsus with 1 basoventral, 1

apical short setae, 1 strong dorsal seta, half as long as basitarsus. Tibia and tarsomere

(from first to fifth) length ratio: fore leg: 87/39/18/14/14/15, mid leg: 116/54/29/

22/17/17, hind leg: 139/53/57/44/26/17.

Wing evenly greyish, slightly fumous on cross veins; costa simple; R 1 reaching to first third of wing; R 2+3 and R 4+5 straight, slightly divergent at apex; ratio of part of costa between R 2+3 and R 4+5 to this between R 4+5 and M 1+2, 29/24; M 1+2 broken in middle of distal part, joining costal vein right before wing tip; R 4+5 and distal part of

M 1 parallel; crossveins m-m and dm-m straight, almost perpendicular to corresponding longitudinal veins; ratio of distal part of M 1 to m-m to distal part of M 1+2 to

dm-m to distal part of M 4, 66/12/44/32/30; anal vein distinct, almost reaching to wing margin; anal lobe well developed; anal angle obtuse; lower calypter yellow,

with black setae; halters yellow.

Abdomen metallic bronze-black, white pollinose laterally, with black hairs and marginal setae; 8th segment black, with sparse black hairs; epandrium black, small,

nearly as long as 5th-6th tergites combined, 2 times longer than high, swollen basally, narrowed distally, concave ventrally, slightly projected distoventrally;

hypandrium basoventral, relatively broad, reaching to distal 4/5 of epandrium,

without teeth; phallus thin and long, with aciculate swelling at base of hypandrium,

with regular sparse row of 7-8 strong ventral teeth behind curvation, gradually decreasing in size towards tip; one minute epandrial seta at base of hypandrium;

distoventral epandrial lobe reduced to 3 long pedunculate setae; surstylus yellow,

bilobate; ventral lobe rounded, with 2 thick spines and few simple setulae; dorsal lobe of surstylus broad, longer than ventral, with membranous apical projection,

with 2 setae and few setulae; postgonite long, half as long as cercus, forked in distal third, with narrow pointed lobes; cercus yellow, widely blackened in distal third,

regularly rounded, dentate distally and ventrally, with double row of straight marginal setae.

MEASUREMENTS (in mm). Body length 2.9; antenna length 0.8; wing length

2.9; wing width 1.0; hypopygium length 0.8.

Female. Unknown.

DISTRIBUTION. Oriental: Indonesia (Bali).

ETYMOLOGY. The species is named after the Russian entomologist Dr. M.V.

Nabozhenko (Rostov-na-Donu).

DIAGNOSIS. The new species is the closest to L. semakau Grootaert et Tang known from Thailand, differing from the latter in evenly greyish wing, which is slightly fumous on cross veins, and in fine structures of hypopygium. L. nabozhenkoi sp. n. cercus is regularly rounded, widely blackened in distal third, without hooked setae, without a strong bristle near the dorsal margin on the inside; phallus with regular sparse row of 7-8 strong ventral teeth behind curvation of phallus,

gradually decreasing in size towards tip. L. semakau cercus is nearly triangular, pale except the black margin, with several hooked bristles and setae distally and with a single strong middorsal bristle on the inside; phallus with about 20 small dense irregular denticles on distal half of ventral side (Tang et al., 2018: Figure 9 ).

Figures

Figs 9–16. Lichtwardtia nabozhenkoi sp. n. 9 – habitus; 10 – head; 11 – antennae; 12 –