Nemastoma bidentatum schmidti Novak, Raspotnig & Slana Novak, 2021

Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor & Raspotnig, Günther, 2021, Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence, European Journal of Taxonomy 777, pp. 1-67 : 30-40

Nemastoma bidentatum schmidti Novak, Raspotnig & Slana Novak ssp. nov.

urn:lsid:zoobank.org:act:389E4C27-24B9-44F9-A724-11475CB9E12B

Figs 2–3 , 4H , 5H , 6H , 7A, H , 8A, H , 9H , 10H , 11H , 12H , 13G ; Table 11

Nemastoma triste – Roewer 1917: 149 [partim: Maribor / Marburg], 151 [partim: Postojna /Adelsberg, Divača / Divaca, Ljubljana / Laibach, Zagreb / Agram].

Nemastoma bidentatum – Roewer 1923: 658 [partim: Postojna /Adelsberg, Ljubljana / Laibach, Zagreb / Agram; the other localities refer to N. bidentatum sparsum ]. — Hadži 1927: 11–14, figs 81−93; 1928: 9; 1931: 108–109 [partim: Podkoren, Peričnik, Bled].

Nemastoma bidentatum bidentatum – Gruber & Martens 1968: 141, figs 3, 23 [partim: Leutscach]. — Marcellino 1973: 194. — Martens 1978: 107, fig 142 [partim: HR: Dugo selo, Zagreb; I: Laghi di

Fusine; SLO: Bled, Cerknica, Ljubljana, Mt. Nanos, Postojna]. — Novak 2004a: 218. Nemastoma quadripunctatum humerale – Hadži 1931: 41, figs 94–98 (by lapsus). Nemastoma (Lugubrostoma) triste pluridentatum – Hadži 1973a: 43 figs 34a–c [partim: Mt. Šmarna

gora, Prestranek].? Nemastoma bidentatum bidentatum × N. bidentatum sparsum – Novak & Gruber 2000: 285.

Diagnosis

Subspecies of Nemastoma bidentatum with elongated oval-trapezoid Ch-Apo, sagittally shallowly bilobe apically, wider distally, with a prominent apical pinnacle posteriorly, and conspicuously club-shaped Pa- Fe, with dorsal margin bent in the middle and nearly straight anterior and posterior portions, and evenly arched ventral margin, with 3 stout spines in distal fourth, and low, inconspicuous Pa-Ti hump. Rec sem of one tubular and 12 slightly elongated vesicles.

Etymology

The subspecies name ʻ schmidti ʼ is dedicated to Ferdinand Joseph Schmidt (1791–1878; Ljubljana), who collected the first specimens of this taxon preserved in his arthropod collection in the Slovenian Museum of Natural History, Ljubljana.

Material examined

Holotype SLOVENIA – VM31 • 1 ♂; Dolenja Sevnica , Poljane nad Škofjo Loko ; 46.12° N, 14.15° E; 517 m a.s.l.; 27 Mar. 2018; T. Novak leg.; mixed forest litter; PMSL-Opiliones-TN 1145/2018.

Paratypes SLOVENIA – VM31 • 2 ♀♀; same collection data as for holotype; PMSL-Opiliones-GR 3809 .

Other material

AUSTRIA – VM74 • 1 ♂; Eisenkappel / Železna Kapla – Ebriach / Obirsko , Carinthia; 7 May 2018; T. Novak leg. (38/2018); PMSL • 1 ♀; Seilach / Sele – Ebriach / Obirsko , Carinthia; 7 May 2018; T. Novak leg. (41/2018); PMSL .

CROATIA – WL67 • 1 ♂, 1 ♀; Mt. Medvednica ; A. Schönhofer leg. (6/2012, rev. 2019); PMSL .

ITALY – UM52 • 7 ♂♂, 7 ♀♀; Artegna ; 4 Mar.−11 Aug. 1992; F. Gasparo leg. (64/2002, rev. 2017); PMSL. – UM62 • 1 ♂; Alta Val Torre ; 5 Aug. 1992; G. Governatori leg. (1062/2003, rev. 2017); PMSL • 1 ♂, 1 ♀; Villanova , Lusevera; 23 Oct. 1997; G. Governatori leg. (244/2003); PMSL • 3 ♂♂, 4 ♀♀; Tarcento / Čenta; 19 Aug. 2001; L. Slana Novak and T. Novak leg. (259/2001, rev. 2017); PMSL • 1 ♂, 5 ♀♀; ibid .; (257/2001, rev. 2018); PMSL • 1 ♂; Valle del Torre Vedronca valley ; 5 Aug. 1992; G. Governatori leg. (1062/2003, rev. 2017); PMSL. – UM65 • 2 ♂♂, 1 ♀; Pontebba ; 19 Aug. 2001; L. Slana Novak and T. Novak leg. (240/2001, rev. 2017); PMSL. – UM80 • 4 ♂♂; Altana / Utana; 26 Aug. 2001; L. Slana Novak and T. Novak leg. (283/2001, rev. 2017); PMSL .

SLOVENIA • 1 ♂, 1 ♀ (Schmidt’s collection); (84/1985 microscopic preparation, JH det. sub N. triste, TN rev.); PMSL • 1 ♂, 1 ♀ (Schmidt’s collection); (89/1985 microscopic preparation, JH det. sub N. triste, TN rev.); PMSL. – UL99 • 1 ♂, 2 ♀♀; Panovec ; 17 Aug. 2000; T. Novak leg. (668/2000, rev. 2017); PMSL • 1 ♀; ibid .; 18 Mar. 2001; S. Brelih leg. (63/2001, TN rev. 2019); PMSL • 9 ♂♂, 5 ♀♀; ibid .; 11 Nov. 2006; L. Slana Novak and T. Novak leg. (152/2006, rev. 2017); PMSL • 1 ♂; Replje ; 8 Apr. 1992; S. Brelih leg. (649/1998, rev. 2017); PMSL. – UM72 • 1 ♂; Mt. Kobariški Stol ; 24 May 2003; L. Slana Novak and T. Novak leg. (720a/2003, TN and SL rev. 2017); PMSL. – UM73 • 1 ♂; Mt. Kobariški Stol ; 8 Jun. 2017; T. Novak leg. (6/2017); PMSL. – UL82 • 1 ♀; Tavenica, Bela ; 10 Oct. 1995; S. Brelih leg. (789/1998, rev. 2018); PMSL • 1 ♂; Sedlo – Mt. Kobariški Stol ; 1 Aug. 1998; L. Slana Novak and T. Novak leg. (52/1999, rev. 2018); PMSL. – UM90 • 2 ♀♀; Avče ; 2 Aug. 2000; T. Novak leg. (298/2000, rev. 2019); PMSL • 2 ♂♂; Doblar ; 2 Aug. 2000; leg. T. Novak (285/2000, rev. 2017); PMSL • 1 ♀; ibid .; 17 Jun. 2001; L. Slana Novak and T. Novak leg. (151/2001, rev. 2017); PMSL • 1 ♂; Lepenka valley, Doblar ; 25 Sep. 1998; L. Slana Novak and T. Novak leg. (1795/1998, rev. 2017); PMSL • 1 ♂; Levpa – Seniški breg ; 2 Aug. 2000; T. Novak leg. (319/2000, rev. 2017); PMSL • 1 ♀; Marija Snežna, Avče ; 2 Aug. 2000; T. Novak leg. (293/2000; rev. 2019); PMSL. – UL91 • 1 ♂; Kamno ; 31 Jul. 1998; L. Slana Novak and T. Novak leg. (1337/1998, rev. 2018); PMSL • 1 ♂, 1 ♀; ibid .; 25 Sep. 1998; L. Slana Novak and T. Novak leg. (1809/1998, rev. 2017); PMSL • 2 ♂♂, 2 ♀♀; Solarji, Livške Ravne ; 8 Aug. 2000; T. Novak leg. (562/2000, TN and SL rev. 2017); PMSL. – UM92 • 6 ♂♂, 1 ♀; Ladrski vrh ; 14 May−11 Jun. 2002; B. Drovenik and A. Gergeli leg. (482/2018); PMSL • 1 ♀; ibid .; 11 Jun.–7 Aug. 2002; B. Drovenik and A. Pirnat leg. (792/2018); PMSL • 2 ♂♂, 3 ♀♀; ibid .; 8 Jul.−7 Aug. 2002; B. Drovenik and A. Pirnat leg. (PK 9/2013, TN rev. 2019); PMSL • 1 ♀; ibid .; (PK 10/2013, TN rev. 2019); PMSL • 1 ♀; ibid .; (585/2018); PMSL • 1 ♀; ibid .; 7 Aug.−4 Sep. 2002; B. Drovenik and A. Pirnat leg. (780/2018); PMSL • 1 ♂; ibid .; (787/2018); PMSL • 2 ♂♂, 6 ♀♀; ibid .; 4 Sep.−1 Oct. 2002; B. Drovenik and A. Pirnat leg. (827/2018); PMSL • 2 ♂♂; ibid .; (910/2018); PMSL • 4 ♀♀; ibid .; (915/2018); PMSL • 3 ♀♀; ibid .; (923/2018); PMSL • 2 ♀♀; ibid .; (928/2018); PMSL • 1 ♂; ibid.; (933/2018); PMSL • 3 ♂♂, 5 ♀♀; ibid .; 1−29 Oct. 2002; B. Drovenik and A. Pirnat leg. (959/2018); PMSL • 1 ♂, 3 ♀♀; ibid .; (961/2018); PMSL • 10 ♀♀; ibid .; (965/2018); PMSL • 4 ♀♀, 1 juv.; ibid .; (971/2018); PMSL • 1 ♂, 1 ♀; ibid .; (974/2018); PMSL • 3 ♂♂, 6 ♀♀; ibid .; (981/2018); PMSL • 8 ♀♀; ibid .; (1003/2018); PMSL • 1 ♂; ibid .; 29 Oct.−7 Nov. 2002; B. Drovenik and A. Gergeli leg. (PK 36/2013, TN rev. 2019); PMSL • 3 ♀♀; ibid .; (PK 37/2013, TN rev. 2019); PMSL • 2 ♂♂, 13 ♀♀; ibid .; (871/2018); PMSL • 2 ♀♀; ibid .; (877/2018); PMSL • 1 ♂, 3 ♀♀; ibid .; (880/2018); PMSL • 2 ♀♀; ibid .; (881/2018); PMSL • 3 ♂♂, 4 ♀♀; ibid .; (897/2018); PMSL • 4 ♂♂, 6 ♀♀; ibid .; (900/2018); PMSL • 3 ♂♂, 1 ♀; ibid .; (939/2018); PMSL • 2 ♀♀; ibid .; 27 Nov.−21 Dec. 2002; B. Drovenik and A. Gergeli leg. (948/2018); PMSL • 3 ♂♂, 2 ♀♀; ibid .; (950/2018); PMSL • 1 ♂; ibid .; (1010/2018); PMSL • 3 ♀♀; ibid .; (1014/2018); PMSL • 2 ♀♀; ibid .; (1019/2018); PMSL • 1 ♂; ibid .; (1026/2018); PMSL • 1 ♂, 2 ♀♀; ibid .; 21 Dec. 2002 − 28 Jan. 2003; B. Drovenik and A. Pirnat leg. (718/2018); PMSL • 1 ♂, 2 ♀♀; ibid .; (742/2018); PMSL • 2 ♂♂, 1 ♀; ibid .; (772/2018); PMSL • 3 ♀♀; ibid .; (778/2018); PMSL • 3 ♂♂, 2 ♀♀; ibid .; (813/2018); PMSL • 2 ♂♂, 3 ♀♀; ibid .; (1007/2018); PMSL • 1 ♂; ibid .; (1013/2018); PMSL. – UL92 • 1 ♂, 1 ♀; near Kamno; Sep.1915; Andreini leg.; PMSL • 1 ♂, 2 ♀♀; Selce ; 10 Oct. 1995; S. Brelih leg. (786/1998, rev. 2017); PMSL • 1 ♂; Šturmi ; 8 Aug. 2000; T. Novak leg. (545/2000, rev. 2017); PMSL. – UL93 • 1 ♀; Bavšica ; 9 Aug. 1996; S. Brelih leg. (744/1998, rev. 2018); PMSL • 3 ♀♀; Brvce ; 11 Oct. 1995; S. Brelih leg. (745/1998, rev. 2018); PMSL. – VL07 • 1 ♀; Velike Žablje ; 17 Aug. 2000; T. Novak leg. (677/2000, rev. 2017); PMSL. – VL09 • 1 ♂, 2 ♀♀; Čepovanska reber ; 2 Aug. 2000; T. Novak leg. (344/2000, rev. 2017); PMSL • 1 ♂; Lijak spring, Ozeljan ; 17 Apr. 2002; B. Drovenik and A. Gergeli leg. (219/2011, rev. 2017); PMSL • 1 ♂; ibid .; 17 Apr.–14 May 2002; B. Drovenik and A. Pirnat leg. (540/2018); PMSL • 2 ♂♂; ibid .; (553/2018); PMSL • 2 ♂♂, 3 ♀♀; ibid .; 14 May−11 Jun. 2002; B. Drovenik and A. Gergeli leg. (503/2018); PMSL • 1 ♀; ibid .; (518/2018); PMSL • 1 ♂; ibid .; 11 Jun. 2002; B. Drovenik and A. Pirnat leg. (212/2011, rev. 2017); PMSL • 1 ♂; ibid .; 8 Jul. 2002; B. Drovenik and A. Pirnat leg. (1093/2018); PMSL • 1 ♀; ibid .; (1097/2018); PMSL • 1 ♀; ibid .; 7 Aug.−4 Sep. 2002; B. Drovenik and A. Gergeli leg. (567/2018); PMSL • 1 ♂; ibid .; 1 Oct. 2002; B. Drovenik and A. Pirnat leg. (313/2018); PMSL • 1 ♂, 1 ♀; ibid .; 18 Feb. 2003; B. Drovenik and A. Gergeli leg. (200/2011, TN rev. 2017); PMSL • 1 ♂; Lokve ; 2 Aug. 2000; T. Novak leg. (350/2000, rev. 2017); PMSL • 1 ♀; ibid .; (364/2000, rev. 2017); PMSL • 1 ♀; Smrečje, Trnovski gozd; 18 Aug. 1973; I. Sivec leg. (519/1981, JG det. 1982, TN rev. 2018); PMSL. – VL16 • Divača – VL19 • 1 ♂; Gorenja Kanomlja ; 1 Aug. 2000; T. Novak leg. (401/2000, rev. 2017); PMSL • 1 ♀; Predmeja ; 21 Aug. 2006; L. Slana Novak and T. Novak leg. (4/2006, rev. 2017); PMSL • 2 ♀♀; Srednja Kanomlja ; 1 Aug. 2000; T. Novak leg. (391/2000, rev. 2017); PMSL. – VL26 • 3 ♂♂, 3 ♀♀; Senožeče ; 20 Sep. 2011; L. Slana Novak and T. Novak leg. (262/2011, rev. 2017); PMSL • 1 ♀; Mt. Vremščica ; 10 Mar. 1994; S. Brelih leg. (680/1998, rev. 2017); PMSL. – VL28 • 2 ♂♂, 1 ♀; Črni vrh – Col; L. Slana Novak and T. Novak leg. (239/2011, rev. 2017); PMSL. – VL29 • 3 ♂♂, 3 ♀♀; Godovič – Črni vrh; 11 Sep. 2011; L. Slana Novak and T. Novak leg. (241/2011, rev. 2017); PMSL • 2 ♂♂, 1 ♀; Gore ; 10 Jun. 2000; L. Slana Novak and T. Novak leg. (155/2000, rev. 2017); PMSL • 1 ♀; Idrija; Jun. 1997; M. Vončina leg. (1702/1998, rev. 2017); PMSL • 1 ♂; ibid .; 10 Jun. 2000; L. Slana Novak and T. Novak leg. (137/2000, rev. 2017); PMSL • 8 ♂♂, 1 ♀; ibid .; (229/2000, TN and SL rev. 2017); PMSL • 1 ♂; ibid .; 1 Aug. 2000; T. Novak leg. (382/2000, rev. 2017); PMSL • 2 ♂♂; Razpotje ; 1 Aug. 2000; T. Novak leg. (408/2000, rev. 2017); PMSL • 1 ♂; Spodnja Idrija ; 10 Jun. 2000; L. Slana Novak and T. Novak leg. (143/2000, rev. 2017); PMSL • 2 ♂♂; Srednja Kanomlja ; 1 Aug. 2000; T. Novak leg. (389/2000, rev. 2017); PMSL • 1 ♀; ibid .; (397/2000, rev. 2017); PMSL • 1 ♂; Urbanovec ; 10 Jun. 2000; L. Slana Novak and T. Novak leg. (149/2000, rev. 2017); PMSL. – VL36 • 1 ♂; Jama Sv. Janeza cave, Cad. No. 897, Prestranek ; 11 Aug. 1952; E. Pretner leg. (56/1983, JH det., TN rev. 2017); PMSL • 1 ♀; ibid .; 29 Aug. 1954; E. Pretner leg. (174/1984, JH det., TN rev. 2018); PMSL • 1 ♂; ibid .; 11 Aug. 1957; E. Pretner leg. (221/1984, TN rev. 2017); PMSL • 3 ♀♀; Jernejeva jama cave , Cad. No. 929, Koče ; 22 Apr. 2001; S. Vidmar, S. Polak, T. Novak leg. (157/2001, rev. 2008); PMSL • Postojna – VL37 • 1 ♂; Belsko ; 14 Apr. 2011; L. Slana Novak and T. Novak leg. (86/2011); PMSL • 1 ♂; Landol ; 9 May 1994; S. Brelih leg. (888/1998); PMSL. – VL38 • 2 ♂♂, 3 ♀♀; Dolenji Novi Svet; L. Slana Novak and T. Novak leg. (230/2011); PMSL • 1 ♀; Novi svet, Hotederšica ; 22 Mar. 1993; S. Brelih leg. (855/1998, rev. 2019); PMSL. – VL39 • 3 ♂♂, 3 ♀♀; Kugu, Rovte ; 11 Sep. 2011; L. Slana Novak and T. Novak leg. (244/2011); PMSL. – VL46 • 1 ♂, 1 ♀; Mt. Javorniki ; 9 May 1994; S. Brelih leg. (836/1998); PMSL • 2 ♂♂, 8 ♀♀; Mali Kožljek, Mt. Javorniki ; 2 Oct. 2011; L. Slana Novak and T. Novak leg. (336/2011); PMSL. – VL47 • 1 ♂, 2 ♀♀; Planina pri Rakeku ; 24 Mar. 1992; F. Velkovrh leg. (221/1995, rev. 2017); PMSL • 1 ♂; Rakov Škocjan ; Jul. 1994; K. Musar leg. (79/1995, rev. 2017); PMSL • 1 ♂; ibid .; 12.–15 May 1994; S. Polak leg. (640/1998, rev. 2018); NM • 7 ♂♂, 17 ♀♀; ibid .; 10 Sep. 2009; A. Schönhofer leg. (3/2012, rev. 2017); PMSL. – VL48 • 1 ♀; Bistra ; 29 Mar. 1993; S. Brelih leg. (641/1998, rev. 2017); PMSL • 1 ♂; Laze ; 16 Oct.−21 Nov. 1994; M. Kuntner leg. (176/1995, rev. 2017); PMSL • 1 ♀; ibid .; Jul.– Aug.1994; M. Kuntner leg. (20/1997, rev. 2017); PMSL • 1 ♂, 1 ♀; ibid .; Aug. 1994; M. Kuntner leg. (183/1995, rev. 2017); PMSL • 1 ♂, 1 ♀; ibid .; Dec. 1994 − Mar. 1995; M. Kuntner leg. (170/1995, rev. 2017); PMSL • 1 ♀; Vranja jama cave , Cad. No. 88, Laze pri Planini ; 7 Jun. 1994; stud. biol. leg. (101/2019); PMSL. – VL49 • 1 ♀; Retovje, Vrhnika ; Jul. 1994, A. Valenčak leg. (17/1997, rev. 2017); PMSL • 1 ♂; ibid .; Jun. 1998; M. Zagmajster leg. (1493/1998, rev. 2017); PMSL • 1 ♀; Vrhnika ; Jul. 1994; A. Valenčak leg. (17/1997, rev. 2017); PMSL. – VL54 • 1 ♂; Mt. Snežnik ; 21 Jul. 2001; B. Bertoncelj, U. Bertoncelj and T. Novak leg. (150/2001, rev. 2017); PMSL • 1 ♀; ibid .; 24 Nov. 2003; S. Polak leg. (120/2006, rev. 2018); NM; PMSL. – VL56 • 1 ♂, 3 ♀♀; Cerkniško jezero ; 19 Oct. 1994; S. Polak leg. (721/1998, rev. 2017); NM • 1 ♀; ibid .; 20 Nov. 1994; S. Polak leg. (687/1998, rev. 2017); NM • 2 ♂♂, 1 ♀; ibid .; 24 Apr. 2001; S. Brelih leg. (72/2001, rev. 2017); PMSL. – VL57 • 2 ♂♂, 4 ♀♀; Dolenje Otave – Selšček ; 14 Aug. 2011; L. Slana Novak and T. Novak leg. (36/2011, rev. 2017); PMSL. – VL58 • 1 ♂; Mt. Krim ; 14 Aug. 2011; L. Slana Novak and T. Novak leg. (42/2011, rev. 2017); PMSL • 2 ♀♀; Rakitna ; 14 Aug. 2011; L. Slana Novak and T. Novak leg. (40/2011, rev. 2017); PMSL. – VL59 • 2 ♂♂, 1 ♀; Bevke ; 20 Oct. 2018; L. Slana Novak and T. Novak leg. (136/2018); PMSL • 1 ♂; Brezovica pri Ljubljani ; 25 May 1997; L. Slana Novak and T. Novak leg. (14/1997, rev. 2018); PMSL • 1 ♀; Draga pri Igu ; 13 Apr. 1995; S. Brelih leg. (732/1998, rev. 2018); PMSL • 3 ♂♂, 10 ♀♀; Žabnica ; 14 Aug. 2011; L. Slana Novak and T. Novak leg. (33/2011, rev. 2017); PMSL. – VL68 • 1 ♂, 3 ♀♀; Iška vas – Mt. Krim ; 14 Aug. 2011; L. Slana Novak and T. Novak leg. (48/2011, rev. 2017); PMSL • 1 ♂; ibid .; 31 Mar. 1997; S. Brelih leg. (53/2001, rev. 2017); PMSL • 1 ♀; Kremenica, Ig ; 29 Jun. 1980 (1155/1981, JG det., TN rev. 2018); PMSL • 1 ♀; ibid .; 2 Mar. 1996; S. Brelih leg. (45/2001, rev. 2017); PMSL • 1 ♂; ibid .; 1 May 1997; S. Brelih leg. (101/2001, rev. 2017); PMSL • 1 ♂; ibid .; 7 Jun. 1997; S. Brelih leg. (117/2001, rev. 2017); PMSL • 1 ♂; ibid .; 10 Jul. 1997; S. Brelih leg. (139/2001, rev. 2017); PMSL • 1 ♂, 1 ♀; ibid .; 25 Aug. 1997; S. Brelih leg. (113/2001, rev. 2017); PMSL • 1 ♀; Mt. Mokrec ; 18 Jul. 1980 (LSN 151/1984, TN rev. 2017); PMSL • 1 ♀; Ščurki, Turjak ; 3 May 2008; L. Slana Novak and T. Novak leg. (155/2011, rev. 2017); PMSL • 1 ♀; ibid .; 20 Jun. 1980; (1149/1981, JG det., TN rev. 2018); PMSL • 1 ♀; Škrilje ; 20 Jun. 1980; (1159/1981, JG det., TN rev. 2018); PMSL. – VL69 • 2 ♀♀; Šmarje Sap ; 10 May 2008; L. Slana Novak and T. Novak leg. (119a/2011, rev. 2017); PMSL • 1 ♂, 1 ♀; ibid .; (147/2011, rev. 2018); PMSL. – VL79 • 1 ♂, 4 ♀♀; Brinjski hrib; L. Slana Novak and T. Novak leg. (99/2012, TN and SL rev. 2017); PMSL • 1 ♂; Stara vas ; 16 Mar. 1993; S. Brelih leg. (665/1998, rev. 2017); PMSL. – VL85 • 1 ♀; Mt. Stojna ; 21 Oct. 1996; S. Brelih leg. (90/2001, rev. 2017); PMSL. – VL89 • 1 ♂, 4 ♀♀; Vintarjevec ; T. Novak leg. (173/2011, rev. 2017); PMSL • 2 ♂♂, 4 ♀♀; ibid .; (178/2011, rev. 2017); PMSL. – VL93 • 1 ♀; Kostel, Dolenja Žaga ; 16 Apr. 1996; S. Brelih leg. (737/1998, rev. 2018); PMSL. – VL99 • 1 ♀; Cerovec, Gabrovka; L. Slana Novak and T. Novak leg. (74/2007, rev. 2017); PMSL. – VM00 • 1 ♂, 2 ♀♀; Cvetrež, Voglarji ; 2 Aug. 2000; T. Novak leg. (324/2000, TN and SL rev. 2017); PMSL • 5 ♂♂, 4 ♀♀; Čepovan ; 1 Aug. 2000; T. Novak leg. (361/2000, rev. 2017); PMSL • 1 ♂; Čepovan – Most na Soči ; 10 Jun. 1997; S. Brelih leg. (110/2001, rev. 2017); PMSL • 1 ♂; Preval, Lokovec, Čepovan ; 2 Aug. 2000; T. Novak leg. (329/2000, rev. 2017); PMSL. – VM05 • several specs; Podkoren ; J. Hadži leg.; PMSL. – VM09 • 1 ♀; Smrečje ; 18 Aug. 1973; I. Sivec leg. (519/1981, JG det., TN rev. 2018); PMSL. – VM10 • 1 ♂; Bukovo ; 8Aug. 2000; T. Novak leg. (501/2000, rev. 2017); PMSL • 3 ♂♂, 5 ♀♀; Oblakov vrh ; 1 Aug. 2000; T. Novak leg. (377/2000, rev. 2017); PMSL • 1 ♂; Stopnik ; 8 Aug. 2000; T. Novak leg. (492/2000, rev. 2018); PMSL. – VM12 • 1 ♀; Šavnica, Bohinjska Bistrica ; 26 Jul. 2002; B. Drovenik and A. Pirnat leg. (1094/2018); PMSL. – VM13 • 1 ♀; Suha ; 25 Oct. 1993; S. Brelih leg. (868/1998, rev. 2018); PMSL. – VM14 • 1 ♀; Peričnik waterfall , Vrata valley ; 14 Aug. 1928; J. Hadži leg.; PMSL. – VM20 • 1 ♀; Fužine ; 8Aug. 2000; T. Novak leg. (484/2000, rev. 2017); PMSL • 1 ♂, 1 ♀; Govejk ; 8 Aug. 2000; T. Novak leg. (486/2000, rev. 2017); PMSL • 1 ♀; Osojnica ; 8 Aug. 2000; T. Novak leg. (498/2000, rev. 2017); PMSL • 2 ♂♂; Otalež ; 8 Aug. 2000; T. Novak leg. (479/2000, rev. 2017); PMSL. – VM21 • 3 ♂♂, 1 ♀; Davča ; 27 Jul. 2003; L. Slana Novak and T. Novak leg. (703/2003, rev. 2017); PMSL • 1 ♀; ibid .; (706/2003, rev. 2017); PMSL • 1 ♂; Velbnik, Mt. Porezen ; 7 Jul. 2003; L. Slana Novak and T. Novak leg. (700/2003); PMSL. – VM22 • 1 ♀; Nomenj ; 25 Oct. 1993; S. Brelih leg. (849/1998, rev. 2018); PMSL. – VM23 • 2 ♀♀; the Pokljuka Mts ; 19 Jul. 1940; J. Hadži leg. (323/1983, rev. 2017); PMSL. – VM30 • 1 ♂, 1 ♀; Gorenja vas – Lučine ; 15 May 1994; S. Brelih leg. (862/1998, rev. 2017); PMSL • 2 ♂♂, 2 ♀♀; Srednja vas – Poljane ; 8 Aug. 2000; T. Novak leg. (483/2000, rev. 2017); PMSL. – VM31 • 15 ♂♂, 14 ♀♀; Dolenja Sevnica, Poljane ; 15 Aug. 2012; L. Slana Novak and T. Novak leg. ( GR 3777−3783 , 3805−3811 , 3819−3825 , 3833−3835 ); PMSL. – VM32 • 1 ♂, 2 ♀♀; Selca ; 15 Aug. 2012; L. Slana Novak and T. Novak leg. (139/2012, TN and SL rev. 2017); PMSL. – VM33 • 1 ♂; Begunje ; 12 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (166/1997, TN and SL rev. 2017); PMSL • 1 ♂; ibid .; (374/1998, rev. 2017); PMSL • Bled. – VM40 • 5 ♂♂, 5 ♀♀; Polhov Gradec – Črni vrh; 11 Sep. 2011; L. Slana Novak and T. Novak leg. (235/2011, rev. 2017); PMSL. – VM41 • 2 ♀♀; Log nad Škofjo Loko ; 8 Aug. 2000; T. Novak leg. (475/2000, rev. 2017); PMSL • 1 ♂; Mt. Lubnik ; Jul. 1998; R. Potočnik leg. (1666/1998, rev. 2017); PMSL • 2 ♂♂, 6 ♀♀; Sopotniška grapa ; 8 Aug. 2000; T. Novak leg. (505/2000, rev. 2017); PMSL • 1 ♂; Štinetova jama cave, Cad. No. 240, Crngrob , Škofja Loka; 25 Jul. 1980; T. Novak, I. Sivec leg. (LSN 214/1984, TN rev. 2019); PMSL. – VM42 • 1 ♂; Bukovščica ; 15 May 1994; S. Brelih leg. (881/1998, rev. 2017); PMSL • 1 ♂; Mt. Šmarjetna gora ; 27 Apr.−4 May 1987; A. Ziherl, T. Trilar leg. (492/1998, rev. 2017); PMSL • 1 ♀; ibid .; 18−24 Oct. 1987; A. Ziherl, T. Trilar leg. (483/1998, rev. 2017); PMSL • 1 ♂; ibid .; (513/1998, TN and SL rev. 2017); PMSL. – VM43 • 1 ♀; Kriška planina; May 1968; (1218/1981, JG det., TN rev. 2017); PMSL • 1 ♂; Križe, Tržič ; 6 Aug. 2015; L. Slana Novak and T. Novak leg. (101/2015, rev. 2017); PMSL. – VM50 • 1 ♀; Ljubljana; Nov. 1986; (631/2002, rev. 2017); PMSL • 1 ♀; ibid .; Nov. 1987; (125/1997, TN and SL rev. 2017); PMSL • 1 ♂, 2 ♀♀; ibid .; Dec. 1987; F. Velkovrh leg. (106/1997, rev. 2017); PMSL • 1 ♂, 1 ♀; ibid .; Mar. 1990; F. Velkovrh leg. (285/1995, rev. 2017); PMSL • 1 ♀; Podutik ; Jun. 1998; (1648/1998, rev. 2017); PMSL • 1 ♀; ibid .; Oct. 1928; J. Staudacher leg. (16/1983 microscopic preparation, rev. 2017); PMSL • 1 ♀; ibid .; May–Jun. 2000; A. Pokrivač leg. (PK and TN 74/2020); PMSL • 1 ♀; Preval , Ljubljana; 18 Mar. 1993; S. Brelih leg. (664/1998, rev. 2017); PMSL. – VM51 • 1 ♂; Mt. Grmada, Medno ; 28 Jul. 1969; (531/1981, rev. 2018); PMSL • 2 ♂♂; ibid .; 3 Jun. 1986; (242/2002, rev. 2017); PMSL • 1 ♂; ibid .; 28 Jul. 1969; (531/1981, rev. 2018); PMSL • 1 ♂; ibid .; 15 May 1970; (526/1981, rev. 2018); PMSL • 1 ♀; ibid .; 19 Apr.−15 May 2002; B. Drovenik and A. Gergeli leg. (188/2011, rev. 2017); PMSL • 1 ♀; ibid .; (446/2018); PMSL • 1 ♀; ibid .; 15 May 2002; B. Drovenik and A. Pirnat leg. (216/2011, rev. 2017); PMSL • 1 ♂; ibid .; 15 May−12 Jun. 2002; B. Drovenik and A. Gergeli leg. (427/2018); PMSL • 1 ♂, 1 ♀; ibid .; (523/2018); PMSL • 1 ♂; ibid .; (558/2018); PMSL • 1 ♀; ibid .; 12 Jun.–9 Jul. 2002; B. Drovenik and A. Gergeli leg. (602/2018); PMSL • 1 ♀; ibid .; (PK 34/2014, TN rev. 2019); PMSL • 2 ♂♂, 1 ♀; ibid .; 8 Aug.–6 Sep. 2002; B. Drovenik and A. Gergeli leg. (PK 86/2012; TN rev. 2019); PMSL • 1 ♂; ibid .; 6 Sep. 2002; B. Drovenik and A. Pirnat leg. (354/2018); PMSL • 1 ♀; ibid .; 2 Oct. 2002 (173/2003, rev. 2017); PMSL • 1 ♀; ibid .; (1100/2018); PMSL • 2 ♀♀; ibid .; (1124/2018); PMSL • 1 ♀; ibid .; 28 Oct.–25 Nov. 2002; B. Drovenik and A. Gergeli leg. (PK 28/2012, TN rev. 2019); PMSL • 1 ♂; ibid .; (264/2018); PMSL • 1 ♀; ibid .; 25 Nov. 2002; B. Drovenik and A. Pirnat leg. (277/2018); PMSL • 4 ♀♀; ibid .; (279/2018); PMSL • 1 ♂; ibid .; 19 Mar. 2003; B. Drovenik and A. Pirnat leg. (330/2018); PMSL • 1 ♀; ibid .; (375/2018); PMSL • 3 ♂♂, 2 ♀♀; Mavčiče ; Jun. 1998; A. Žunič leg. (1568/1998, rev. 2017); PMSL. – VM52 • 14 ♂♂, 12 ♀♀; Predoslje ; 29 Jan. 1954; A. Polenec leg. (263/1984, rev. 2017); PMSL • 11 ♂♂, 23 ♀♀; ibid .; 31 Mar. 1954; A. Polenec leg. (337/1984, rev. 2017); PMSL • 8 ♂♂, 7 ♀♀; ibid .; 10 Oct. 1954; A. Polenec leg. (106/1984, rev. 2017); PMSL • 3 ♂♂, 2 ♀♀; ibid .; 15 Oct. 1954; A. Polenec leg. (351/1984, TN and SL rev. 2017); PMSL • 6 ♂♂, 14 ♀♀; ibid .; 24 Oct. 1954, A. Polenec leg. (63/1984, rev. 2017); PMSL • 4 ♂♂, 7 ♀♀; ibid .; 30 Oct. 1954; A. Polenec leg. (416/1984, TN and SL rev. 2017); PMSL • 2 ♂♂, 4 ♀♀; ibid .; 6 Nov. 1954; A. Polenec leg. (556/1983, TN and SL rev. 2017); PMSL • 5 ♂♂, 11 ♀♀; ibid .; (319/1984, rev. 2017); PMSL • 2 ♂♂; ibid .; 14 Nov. 1954; A. Polenec leg. (345/1984, TN and SL rev. 2017); PMSL • 3 ♂♂, 9 ♀♀; ibid .; (393/1984, TN and SL rev. 2017); PMSL • 3 ♂♂, 1 ♀; ibid .; 28 Nov. 1954; A. Polenec leg. (364/1984, TN and SL rev. 2017); PMSL • 1 ♂, 4 ♀♀; ibid .; (410/1984, TN and SL rev. 2017); PMSL • 28 ♂♂, 32 ♀♀; ibid .; 5 Dec. 1954; A. Polenec leg. (382/1984, TN and SL rev. 2017); PMSL • 1 ♀; ibid .; 18 Dec. 1954; A. Polenec leg. (20/1984, rev. 2017); PMSL • 32 ♂♂, 34 ♀♀; ibid .; (58/1984, rev. 2017); PMSL • 1 ♀; ibid .; 31 Dec. 1954; A. Polenec leg. (24/1984, rev. 2017); PMSL • 9 ♂♂, 21 ♀; ibid .; 9 Jan. 1955; A. Polenec leg. (324/1984, TN and SL rev. 2017); PMSL • 12 ♂♂, 8 ♀♀; ibid .; 15 Jan. 1955; A. Polenec leg. (209/1984, TN and SL rev. 2017); PMSL • 1 ♂; ibid .; (210/1984, TN and SL rev. 2017); PMSL • 6 ♂♂, 17 ♀♀; ibid .; 22 Jan. 1955; A. Polenec leg. (342/1984, TN and SL rev. 2017); PMSL • 18 ♂♂, 15 ♀♀; ibid .; 7 Feb. 1955; A. Polenec leg. (103/1984, TN and SL rev. 2017); PMSL • 20 ♂, 27 ♀♀; ibid .; 12 Feb. 1955; A. Polenec leg. (242/1984, TN and SL rev. 2017); PMSL • 1 ♀; ibid .; 15 Feb. 1955; A. Polenec leg. (328/1984, TN and SL rev. 2017); PMSL • 7 ♂♂, 14 ♀♀; ibid .; 19 Feb. 1955; A. Polenec leg. (390/1984, TN and SL rev. 2017); PMSL • 2 ♀♀; ibid .; 25 Mar. 1955; A. Polenec leg. (29/1984, TN and SL rev. 2017); PMSL • 1 ♂; ibid .; (33/1984, TN and SL rev. 2017); PMSL • 7 ♂♂, 7 ♀♀; ibid .; 27 Mar. 1955; A. Polenec leg. (285/1984, TN and SL rev. 2017); PMSL • 8 ♂♂, 12 ♀♀; ibid .; 29 Mar. 1955; A. Polenec leg. (99/1984, TN and SL rev. 2017); PMSL • 9 ♂♂, 10 ♀♀; ibid .; (277/1984, TN and SL rev. 2017); PMSL • 2 ♂♂, 2 ♀♀; ibid.; 6 Apr. 1955; A. Polenec leg. (90/1984); PMSL • 2 ♂♂, 2 ♀♀; ibid.; 9 Apr. 1955; A. Polenec leg. (229/1984, TN and SL rev. 2017); PMSL • 1 ♂, 1 ♀; ibid.; 8 May 1955; A. Polenec leg. (69/1984, TN and SL rev. 2017); PMSL • 4 ♂♂, 6 ♀♀; ibid.; (247/1984, TN and SL rev. 2017); PMSL • 7 ♂♂, 5 ♀♀; ibid.; 21 May 1955; A. Polenec leg. (279/1984, TN and SL rev. 2017); PMSL • 9 ♂♂, 3 ♀♀; ibid.; 4 Jun. 1955; A. Polenec leg. (298/1984, TN and SL rev. 2017); PMSL • 1 ♀; ibid.; (333/1984, TN and SL rev. 2017); PMSL • 2 ♀♀; ibid.; 23 Apr. 1955; A. Polenec leg. (313/1984, TN and SL rev. 2017); PMSL • 13 ♂♂, 11 ♀♀; ibid.; 21 Jun. 1955; A. Polenec leg. (50/1984, TN and SL rev. 2017); PMSL • 3 ♀♀; ibid.; 2 Jul. 1955 (379/1984, TN and SL rev. 2017); PMSL • 1 ♀; ibid.; 27 Aug. 1955; A. Polenec leg. (253/1984, TN and SL rev. 2017); PMSL • 1 ♀; ibid.; (305/1984, TN and SL rev. 2017); PMSL • 2 ♀♀; ibid.; 8 Sep. 1955; A. Polenec leg. (258/1984, TN and SL rev. 2017); PMSL • 7 ♂♂, 12 ♀♀; ibid.; 8 Oct. 1955; A. Polenec leg. (362/1983, TN and SL rev. 2017); PMSL • 1 ♂, 2 ♀♀; ibid.; 22 Oct. 1955; A. Polenec leg. (383/1983, TN and SL rev. 2017); PMSL • 3 ♂♂, 4 ♀♀; ibid.; (389/1983, rev. 2017); PMSL • 11 ♂♂, 9 ♀♀; ibid.; 9 Nov. 1955; A. Polenec leg. (91/1984, TN and SL rev. 2017); PMSL • 2 ♂♂, 3 ♀♀; ibid.; 16 Apr. 1956; A. Polenec leg. (482/1983, TN and SL rev. 2017); PMSL • 1 ♂, 2 ♀♀; ibid.; 22 Apr. 1956; A. Polenec leg. (280/1983, TN and SL rev. 2017); PMSL • 1 ♀; ibid.; 26 May 1957; A. Polenec leg. (441/1983, TN and SL rev. 2017); PMSL • 1 ♂; ibid.; 4 Jun. 1956; A. Polenec leg. (289/1984, TN and SL rev. 2017); PMSL • 1 ♀; ibid.; 22 Jun. 1956; A. Polenec leg. (472/1983); PMSL • 1 ♂; ibid.; 8 Jul. 1956; A. Polenec leg. (208/1983, TN and SL rev. 2017); PMSL • 1 ♂; ibid.; 22 Jul. 1956; A. Polenec leg. (LSN 9/1983, TN rev. 2018); PMSL • 1 ♂; ibid.; (532/1983, TN and SL rev. 2017); PMSL • 1 ♂; ibid.; 10 Aug. 1956; A. Polenec leg. (256/1983, TN and SL rev. 2017); PMSL • 1 ♂, 1 ♀; ibid.; 24 Oct. 1956; A. Polenec leg. (537/1983, TN and SL rev. 2017); PMSL • 5 ♂♂, 11 ♀♀; ibid.; 29 Oct. 1956; A. Polenec leg. (508/1983, TN and SL rev. 2017); PMSL • 5 ♂♂, 1 ♀; ibid.; 30 Nov. 1956; A. Polenec leg. (423/1983, TN and SL rev. 2017); PMSL • 1 ♂, 5 ♀♀; ibid.; 16 Dec. 1956; A. Polenec leg. (196/1983, TN and SL rev. 2017); PMSL • 7 ♂♂, 11 ♀♀; ibid.; (544/1983, TN and SL rev. 2017); PMSL • 2 ♂♂, 2 ♀♀; ibid.; 30 Dec. 1956; A. Polenec leg. (214/1983, TN and SL rev. 2017); PMSL • 4 ♀♀; ibid.; (295/1984, TN and SL rev. 2017); PMSL • 8 ♂♂, 10 ♀♀; ibid.; 20 Feb. 1957; A. Polenec leg. (449/1983, TN and SL rev. 2017); PMSL • 4 ♂♂, 10 ♀♀; ibid.; 23 Feb. 1957; A. Polenec leg. (418/1983, TN and SL rev. 2017); PMSL • 5 ♂♂, 5 ♀♀; ibid.; (215/1984, JH det., TN and SL rev. 2017); PMSL • 4 ♂♂, 10 ♀♀; ibid.; 8 Mar. 1957; A. Polenec leg. (247/1983, TN and SL rev. 2017); PMSL • 5 ♂♂, 12 ♀♀; ibid.; 6 Apr. 1957; A. Polenec leg. (491/1983, TN and SL rev. 2017); PMSL • 10 ♂♂, 11 ♀♀; ibid.; 9 Mar. 1957; A. Polenec leg. (465/1983, TN and SL rev. 2017); PMSL • 12 ♂♂, 4 ♀♀; ibid.; 3 May 1957; A. Polenec leg. (506/1983, TN and SL rev. 2017); PMSL • 5 ♂♂, 5 ♀♀; ibid.; 18 May 1957; A. Polenec leg. (551/1983, TN and SL rev. 2017); PMSL. – VM60 • 12 ♂♂, 29 ♀♀; Ljubljana; Jun. 1972; (518/1981, JG det., TN rev. 2020); PMSL • 1 ♂; ibid.; 26 May 1986; Hribernik, Koren and Mencinger leg. (255/2002, rev. 2017); PMSL. – VM61 • 4 ♂♂, 1 ♀; Mt. Gobavica , Mengeš; 22 Aug. 2011; L. Slana Novak and T. Novak leg. (97/2011, rev. 2017); PMSL • 1 ♀; Rašica ; Jun. 1994; H. Urbas leg. (241/1995, TN rev. 2017); PMSL. – VM62 • 1 ♂, 1 ♀; Stahovica – Ambrož ; 17 Aug. 2011; L. Slana Novak and T. Novak leg. (61/2011, rev. 2017); PMSL. – VM63 • 1 ♀; Mt. Kalški greben ; 12 Aug. 2009; L. Slana Novak and T. Novak leg. (28/2009, rev. 2017); PMSL • 1 juv.; ibid.; (31/2009, rev. 2017); PMSL • 1 ♂, 2 ♀♀; Kamnik; 29 Oct. 2000; N. Žmaher leg. (885/2000, rev. 2017); PMSL • 1 ♀; Mt. Koren ; 8 Sep. 2009; L. Slana Novak and T. Novak leg. (50/2009, rev. 2017); PMSL • 1 ♂; Mt. Košutna ; 22 Sep. 2009; L. Slana Novak and T. Novak leg. (68/2009, rev. 2017); PMSL • 1 ♀; Mt. Mokrica ; 22 Sep. 2009; L. Slana Novak and T. Novak leg. (80/2009, rev. 2017); PMSL. – VM70 • 1 ♂, 3 ♀♀; Jevnica ; 8 Jul. 1986; L. Slana Novak and T. Novak leg. (538/2002, rev. 2017); PMSL • 5 ♂♂, 3 ♀♀; Senožeti – Velika vas; 17 Aug. 2011; L. Slana Novak and T. Novak leg. (55/2011, rev. 2017); PMSL • 2 ♂♂, 3 ♀♀; Spodnja Javoršica ; 16 Aug. 2000; T. Novak leg. (603/2000, rev. 2017); PMSL • 1 ♂; Vinje ; 16 Aug. 2000; T. Novak leg. (693/2000, rev. 2017); PMSL. – VM71 • 1 ♂; Brezje pri Dobu ; 16 Aug. 2000; T. Novak leg. (605/2000, rev. 2017); PMSL • 5 ♂♂, 1 juv.; Lukovica pri Domžalah; L. Slana Novak and T. Novak leg. (64/2011, rev. 2017); PMSL • 1 ♀; Prevoje pri Šentvidu , Šentvid pri Lukovici ; 30 Mar. 1993; S. Brelih leg. (637/1998, rev. 2018); PMSL. – VM72 • 7 ♂♂; Smrečje v Črni ; L. Slana Novak and T. Novak leg. (73/2011, rev. 2017); PMSL. – VM73 • 1 ♂, 1 ♀; Robanov kot ; 4 Aug. 2015; T. Novak leg. (85/2015, rev. 2017); PMSL. – VM74 • 2 ♀♀; Podolševa ; 4 Aug. 2015; T. Novak leg. (83/2015, rev. 2017); PMSL. – VM80 • 3 ♂♂, 3 ♀♀; Kresnice − Litija; 9 Jul. 1986; L. Slana Novak and T. Novak leg. (496/2002, rev. 2017); PMSL • 1 ♀; Ribče ; 20 Nov. 1992; S. Brelih leg. (724/1998, rev. 2017); PMSL. – VM81 • 6 ♂♂, 4 ♀♀; Mt. Limbarska gora ; 17 Aug. 2011; L. Slana Novak and T. Novak leg. (68/2011, rev. 2017); PMSL • 1 ♀; Negastern ; 16 Aug. 2000; T. Novak leg. (593/2000, rev. 2017); PMSL • 1 ♀; Podmilj ; 10 May 2008; L. Slana Novak and T. Novak leg. (106/2012, rev. 2017); PMSL. – VM82 • 3 ♂♂, 1 ♀; Mt. Gradišče, Bočna , Gornji Grad; 17 Aug. 2011; L. Slana Novak and T. Novak leg. (75/2011, rev. 2017); PMSL. – VM83 • 1 ♂; Primož pri Ljubnem ; 31 Mar. 1994; S. Brelih leg. (875/1998, rev. 2017); PMSL. – VM84 • 1 ♀; Bistra ; 4 Aug. 2015; T. Novak leg. (89/2015, rev. 2017); PMSL. – VM85 • 1 ♂, 1 ♀; Najbrževo, Mt. Peca ; 10 Aug. 1999; F. Janžekovič leg. (82/2002, JG det., TN rev. 2018); PMSL • 1 ♂; ibid.; 21 Oct. 1999; F. Janžekovič leg. (45/2003, rev. 2018); PMSL • 1 ♂, 2 ♀♀; Topla valley ; 12 Jul. 2003; L. Slana Novak and T. Novak leg. (187/2003, rev. 2017); PMSL. – VM90 • 1 ♀; Cerovec, Gabrovka ; 15 Aug. 2007; L. Slana Novak and T. Novak leg. (74/2007, rev. 2017); PMSL • 2 ♂♂, 2 ♀♀; Renke ; 9 Jul. 1986; L. Slana Novak and T. Novak leg. (518/2002, rev. 2017); PMSL • 3 ♂♂, 2 ♀♀; Sava ; 15 Aug. 2007; L. Slana Novak and T. Novak leg. (55/2007, rev. 2017); PMSL. – VM91 • 1 ♂, 4 ♀♀; Mt. Čemšenik ; 1 Jul. 2002; L. Slana Novak and T. Novak leg. (1202/2002, rev. 2018); PMSL • 5 ♂♂, 1 ♀; Mt. Črni vrh, Mt. Čemšeniška planina ; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1171/2002, rev. 2018); PMSL • 1 ♀; Sv. Primož in Felicijan, Mt. Čemšeniška planina ; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1197/2002, rev. 2017); PMSL. – VM93 • 1 ♂, 1 ♀; Mt. Golte ; 14 Dec. 1984; A. Polenec leg. (LSN 57/1985, TN rev. 2018); PMSL • 1 ♂; Žlabor ; 31 Mar. 1994; S. Brelih leg. (886/1998, rev. 2017); PMSL. – VM94 • 1 ♂, 1 ♀; Florjan ; 6 Jul. 2003; L. Slana Novak and T. Novak leg. (196/2003, rev. 2018); PMSL • 1 ♂; Mt. Smrekovec ; 18 Sep. 2000; (579/2002, rev. 2017); PMSL. – VM95 • 3 ♂♂; Tolsti vrh , Ravne na Koroškem; 11 Oct. 2006; T. Novak leg. (55/2006, rev. 2019); PMSL • 4 ♂♂, 3 ♀♀; Mt. Uršlja gora; L. Slana Novak and T. Novak leg. (65/2006, TN and SL rev. 2017); PMSL. – VM96 • 2 ♂♂, 3 ♀♀; Jamnica ; 10 Oct. 2006; T. Novak leg. (11/2006, rev. 2019); PMSL • 1 ♂, 1 ♀; Zgornja Jamnica ; Ravne na Koroškem; 10 Oct. 2006; T. Novak leg. (13/2006, rev. 2018); PMSL. – WL07 • 1 ♂; Mali Lipovec ; 10 May 2008; L. Slana Novak and T. Novak leg. (107/2011, rev. 2017); PMSL. – WL19 • 1 ♂; Krmelj – Tržišče ; 15 Aug. 2007; L. Slana Novak and T. Novak leg. (66/2007, rev. 2017); PMSL. – WL39 • 1 ♂; Arto ; 18 Nov. 1994; S. Brelih leg. (188/2001, rev. 2017); PMSL. – WL48 • 1 ♂, 4 ♀♀; Prilipe , Brežice; 26 May 1984; L. Slana Novak and T. Novak leg. (LSN 228/1984, TN rev. 2018); PMSL. – WM00 • 1 ♀; Sopota valley, Podkum ; 1 Apr. −6 May 2002; B. Drovenik and A. Pirnat leg. (642/2018); PMSL • 1 ♀; ibid.; (648/2018); PMSL • 1 ♂; ibid.; 27 May 2002; B. Drovenik and A. Pirnat leg. (214/2011, ver. 2017); PMSL • 1 ♂; ibid.; 27 May−1 Jul. 2002; B. Drovenik and A. Gergeli leg. (185/2011, rev. 2017); PMSL • 3 ♀♀; ibid.; (187/2011, rev. 2017); PMSL • 1 ♀; ibid.; (189/2011, rev. 2017); PMSL • 2 ♀♀; ibid.; 20 Aug.–19 Sep. 2002; B. Drovenik and A. Gergeli leg. (PK 24/2012, TN rev. 2018); PMSL • 1 ♀; ibid.; (858/2018); PMSL • 1 ♂, 2 ♀♀; ibid.; (888/2018); PMSL • 1 ♂; ibid.; 19. Sep. 2002; B. Drovenik and A. Pirnat leg. (256/2018); PMSL • 1 ♀; ibid.; 19 Sep.−15 Oct. 2002; B. Drovenik and A. Pirnat leg. (943/2018); PMSL • 1 ♀; ibid.; (985/2018); PMSL • 1 ♀; ibid.; 15 Oct. 2002; B. Drovenik and A. Pirnat leg. (1030/2018); PMSL • 1 ♀; ibid.; (1078/2018); PMSL • 2 ♀♀; ibid.; 15 Oct. 2002 – 12 Nov. 2002; B. Drovenik and A. Gergeli leg. (PK 81/2012, TN rev. 2018); PMSL • 1 ♂; ibid.; (898/2018); PMSL • 3 ♂♂, 2 ♀♀; ibid.; 12 Nov. 2002 − 6 Mar. 2003; B. Drovenik and A. Pirnat leg. (685/2018); PMSL • 1 ♂; ibid.; (713/2018); PMSL • 1 ♀, 1 juv.; ibid.; (736/2018); PMSL • 1 ♂, 3 ♀♀; ibid.; (743/2018); PMSL • 1 juv.; ibid.; 6 Mar. 2003; B. Drovenik and A. Pirnat leg. (212/2018); PMSL • 1 ♀; ibid.; (238/2018); PMSL • 1 ♂; Šklendrovec, Mt. Kum ; 22 Apr. 1992; S. Brelih leg. (727/1998, rev. 2017); PMSL. – WM02 • 3 ♂♂, 1 ♀; Braslovško jezero , Braslovče; 26 Oct. 2013; T. Novak leg. (35/2013, rev. 2017); PMSL. – WM03 • 3 ♂♂, 3 ♀♀; Andraž nad Polzelo ; 26 Oct. 2013; T. Novak leg. (47/2013, rev. 2017); PMSL. – WM04 • 6 ♂♂; Mislinjska Dobrava ; 26 Oct. 2013; T. Novak leg. (37/2013, rev. 2018); PMSL • 1 ♀; Rdečka jama cave, Cad. No. 3488, Mt. Rdečki vrh ; 10 Jul. 1977; V. Kuštor, T. Novak leg. (1119/1981, JG det., TN rev. 2018); PMSL. – WM06 • 1 ♂, 1 ♀; Dom na Košenjaku ; 14 Oct. 1995; S. Hudrap and M. Pavlin leg. (LSN 103/1996, TN rev. 2004); PMSL • 1 ♂, 1 ♀; Mt. Košenjak ; 14 Oct. 1995; S. Hudrap and M. Pavlin leg. (LSN 103/1996, TN rev. 2019); PMSL • 2 ♂♂; Pernice ; 18 Aug. 2005; T. Novak leg. (57/2005, rev. 2018); PMSL • 1 ♂; ibid.; 18 Aug. 2005; T. Novak leg. (66a/2005, rev. 2017); PMSL • 1 ♂; ibid.; 1 Aug. 2005; T. Novak leg. (69/2005, rev. 2019); PMSL • 2 ♂♂, 3 ♀♀; ibid.; 20 Aug. 2005; L. Slana Novak and T. Novak leg. (86/2005, rev. 2017); PMSL • 1 ♂; ibid.; 22 Aug. 2005; L. Slana Novak and T. Novak leg. (108/2005, rev. 2017); PMSL • 1 ♂; ibid.; (110/2005, rev. 2017); PMSL • 1 ♀; ibid.; (112/2005, rev. 2017); PMSL • 2 ♂♂, 1 ♀; ibid.; (114/2005, rev. 2018); PMSL. – WM13 • 1 ♂, 1 ♀; Lipje, Vinska Gora ; 21 Apr. 2002; L. Slana Novak and T. Novak leg. (741/2002, rev. 2018); PMSL • 1 ♂, 1 ♀; Vodosteč, Črnova ; 21 Apr. 2002; L. Slana Novak and T. Novak leg. (737/2002, rev. 2019); PMSL • 1 ♂, 1 ♀; Zavrh nad Dobrno ; 22 Apr. 1995; S. Brelih leg. (770/1998, rev. 2018); PMSL. – WM14 • 2 ♀♀; Paka pri Velenju ; 9 Jun. 2015; T. Novak leg. (50/2015, rev. 2018); PMSL. – WM15 • 3 ♂♂, 1 ♀; Sv. Primož ; 22 Oct. 1995; S. Hudrap and M. Pavlin leg. (LSN 104/1996, TN rev. 2018); PMSL • 1 ♂, 1 ♀; Mt. Kremžarjev vrh ; 26 May 2005; L. Slana Novak and T. Novak leg. (187/2005, rev. 2017); PMSL. – WM16 • 2 ♂♂; Mt. Kapunar ; 20 Aug. 2005; L. Slana Novak and T. Novak leg. (60/2005, rev. 2018); PMSL • 1 ♂; Pernice ; 20 Aug. 2005; L. Slana Novak and T. Novak leg. (75/2005, rev. 2018); PMSL • 1 ♀; ibid.; 27 Apr. 2008; L. Slana Novak and T. Novak leg. (167/2011, rev. 2017); PMSL • 1 ♀; Prevalov graben , Muta; 27 Apr. 2008; L. Slana Novak and T. Novak leg. (150/2011, rev. 2017); PMSL • 1 ♀; Radlje ob Dravi; 2 Jul. 1939; (334/1983, rev. 2017); PMSL • 1 ♀; ibid.; 15 May 1966; I. Leitinger leg. (510/1981, rev. 2018); PMSL • 1 ♀; ibid.; (513/1981, JG det., TN rev. 2018); PMSL • 1 ♂; ibid.; 12 Jun. 1966; I. Leitinger leg. (514/1981, rev. 2018); PMSL • 1 ♂; ibid.; (517/1981, rev. 2018); PMSL • 1 ♀; ibid.; 10 Jul. 1966; I. Leitinger leg. (525/1981, JG det., TN rev. 2018); PMSL • 1 ♀; ibid.; 8 Aug. 1966; I. Leitinger leg. (1222/1981, JG det., TN rev. 2018); PMSL • 3 ♂♂; ibid.; 26 Aug. 1966; I. Leitinger leg. (511/1981, rev. 2018); PMSL • 1 ♀; ibid.; (512/1981, JG det., TN rev. 2018); PMSL • 2 ♀♀; ibid.; (515/1981, JG det., TN rev. 2018); PMSL • 1 ♀; ibid.; (524/1981, JG det., TN rev. 2018); PMSL • 1 ♀; ibid.; 9 Sep. 1966; I. Leitinger leg. (516/1981, JG det., TN rev. 2018); PMSL • 1 ♀; ibid.; (671/1981, rev. 2017); PMSL • 1 ♀; ibid.; (152/1982, JG det., TN rev. 2018); PMSL • 1 ♀; ibid.; 8 Oct. 1973; T. Novak leg. (994/1981, rev. 2018); PMSL • 3 ♂♂, 1 ♀; Sv. Trije Kralji ; 19 Aug. 2005; T. Novak leg. (97/2005, rev. 2017); PMSL • 1 ♀; ibid.; 20 Aug. 2005; L. Slana Novak and T. Novak leg. (82/2005, rev. 2017); PMSL • 2 ♂♂; ibid.; (73/2005, rev. 2018); PMSL. – WM23 • 2 ♂♂, 1 ♀; Mt. Stenica ; 25 Apr. 1982; T. Novak; G. Kolar leg. (267/1982, JG det., TN rev. 2018); PMSL. – WM32 • 1 ♂; Razbor, Ponikva ; 9 Jun. 2015; T. Novak leg. (54/2015, rev. 2017); PMSL. – WM33 • 6 ♂♂, 7 ♀♀; Preloge ; 17 Aug. 1983; L. Slana Novak and T. Novak leg. (LSN 73/1983, TN rev. 2018); PMSL. – WM34 • 1 ♀; Sv. Trije Kralji, the Pohorje Mts ; 10 Aug. 1984; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 250/1984, TN rev. 2018); PMSL. – WM35 • 1 ♀; Smolnik ; 8 Aug. 2015; L. Slana Novak and T. Novak leg. (108/2015, TN rev. 2017); PMSL. – WM36 • 2 ♂♂, 2 ♀♀; Ožbalt ; 25 Oct. 2013; T. Novak leg. (45/2013, rev. 2017); PMSL. – WM43 • 1 ♀; Brezje pri Poljčanah ; 10 Apr. 1995; S. Brelih leg. (714/1998, rev. 2017); PMSL • 1 ♀; Zbelovo ; 13 Jul. 2002; L. Slana Novak and T. Novak leg. (1158/2002, rev. 2018); PMSL. – WM45 • 1 ♀; Sv. Bolfenk, the Pohorje Mts ; 27 Jul. 1987; F. Janžekovič leg. (5/1999, TN rev. 2018); PMSL • 1 ♂; Limbuš ; 11 Oct. 2001; D. Kos leg. (217/2004, rev. 2017); PMSL • 16 ♂♂, 36 ♀♀; ibid.; (224/2004, rev. 2017); PMSL • 1 ♂; ibid.; (218/2004, rev. 2018); PMSL • 7 ♂♂; ibid.; (225/2004, rev. 2018); PMSL. – WM46 • 1 ♀; Gaj nad Mariborom ; 16 Apr. 1983; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 64/1983, TN rev. 2018); PMSL • 1 ♀; ibid.; 20 Apr. 2008; L. Slana Novak and T. Novak leg. (164/2011, rev. 2018); PMSL • 1 juv.; Gaj nad Mariborom − Žavcarjev vrh ; 1 Aug. 1981; T. Novak leg. (1084/1981, rev. 2019); PMSL • 1 ♀; Jurski vrh ; 20 Apr. 2008; L. Slana Novak and T. Novak leg. (166/2011, rev. 2017); PMSL • 2 ♂♂, 2 ♀♀; Kamniška graba ; 20 Apr. 2008; L. Slana Novak and T. Novak leg. (129/2011, rev. 2018); PMSL • 5 ♂♂, 1 ♀; Zgornji Slemen ; 16 Apr. 1983; T. Novak, M. Slana Novak, L. Slana Novak and M. Štangelj leg. (LSN 10/1983, TN rev. 2018); PMSL. – WM54 • 1 ♂, 1 ♀; Marjeta na Dravskem polju ; 8 Jun. 2015; P. Kozel leg. (57/2015, rev. 2017); PMSL. – WM55 • 1 ♀; Maribor – Ruperče ; 20 Apr. 1993; S. Brelih leg. (852/1998, rev 2018); PMSL. – WM64 • 1 ♂; Krčevina pri Vurbergu ; 15 Sep. 2015; T. Marinko leg. (134/2015, TN rev. 2017); PMSL • 1 ♂; Placar ; 16 Oct. 1994; M. Varžič leg. (17/1999, tev. 2018); PMSL. – WM65 • 1 ♂; Hum ; 23 Apr. 2008; L. Slana Novak and T. Novak leg (130/2011, rev. 2017); PMSL • 2 ♂♂, 2 ♀♀; Zgornja Voličina – Hrastovec ; 2 Oct. 2002; T. Novak leg. (1284/2002, rev. 2018); PMSL. – WM74 • 1 ♂; Hlaponci ; 23 Apr. 1998; S. Brelih leg. (73/2001, rev. 2017); PMSL • 1 ♂; ibid.; (74/2001, rev. 2017); PMSL • 3 ♂♂, 2 ♀♀; Pacinje ; 13 Aug. 1983; T. Novak and L. Slana Novak leg. (LSN 60/1983, T. Novak rev. 2018); PMSL. – WM75 • 2 ♀♀; Drbetinci ; 28 Apr. 2002; L. Slana Novak and T. Novak leg. (731/2002, rev. 2017); PMSL • 1 ♂, 1 ♀; Kokolanjšček ; 26 Sep. 2005; L. Slana Novak and T. Novak leg. (121/2005, rev. 2017); PMSL • 1 ♂, 1 ♀; Velika Slavšina ; 13 Oct. 2005; L. Slana Novak and T. Novak leg. (149/2005, rev. 2017); PMSL • 3 ♂♂; Vitomarci ; 2 Jun. 2002; L. Slana Novak and T. Novak leg. (756/2002, rev. 2018); PMSL. – WM76 • 1 ♂; Kočki vrh ; 30 Jun. 2002; T. Novak leg. (784/2002, rev. 2017); PMSL. – WM95 • 1 ♂; Gresovščak; L. Slana Novak and T. Novak leg. (162/2012, TN and SL rev. 2017); PMSL .

Description

Male (holotype)

BODY. Length 1.94, width 1.43.

CHELICERAE. Ch basal article (without Apo) length 0.58, 2.0 times as long as wide at Ch max width at dorsal hump. Ch-Apo elongated oval-trapezoid, shallowly bilobe apically, wider distally, with a prominent apical pinnacle posteriory. Ch-Apo height 0.21, min width 0.08 (at the base) and max width 0.15. Secretion field frontally. Distal article length 0.60, max width 0.25, movable finger length 0.28 ( Figs 4H , 6H , 9H ).

PEDIPALPS. Pa-Tr with dorsal margin conspicuously inclined distally. Pa-Fe conspicuously club-shaped (Pa-Fe min:max width ~1:2.8) with dorsal margin bent in the middle and nearly straight anterior and posterior portions, and evenly arched ventral margin, with 3 stout spines in distal fourth. Pa-Ti hump low, inconspicuous ( Figs 5H , 6H ). Pa-Pt length:max width ~1:4.1. Pa article lengths in Table 11.

PENIS. Pe length 1.47, base 0.48, glans 0.09, stylus 0.07 ( Figs.7A, H , 8H ).

LEGS. Pseudoarticle leg-Fe formula I−IV: 0−1−1−2. Tarsomere leg-Fe formula I−IV: 10−17−10−10. Leg article lengths in Table 11.

Female

BODY. Length 2.11, width 1.65.

CHELICERAE. Ch basal article width 0.63, 2.7 times as long as wide at dorsal hump, with nearly straight dorsal margin in front of indentation and ventral margin ( Fig. 13G ). Distal article length 0.64, max width 0.23, movable finger length 0.27.

PEDIPALPS. Pa-Tr with low dorsal margin, highest in the first third and straight posteriorly. Pa-Fe with conspicuous distal bonce, Pa-Pt, Pa-Ti and Pa-Ta elongated ( Fig. 13G ). Pa article lengths in Table 11.

OVIPOSITOR. Ovipositor length 0.84, Rec sem of one tubular and 12 slightly elongated vesicles ( Fig. 11G ).

LEGS. Leg article lengths in Table 11.

Remarks

Hadži (1927, sub N. bidentatum ) made the first detailed drawings of N. b. schmidti ssp. nov. from Schmidt’s collection. Prior to this study, all published evidence citing either N. bidentatum or N. b. bidentatum in Slovenia (all localities), Croatia (Zagreb) and NE Italy (all localities) refers to N. b. schmidti ssp. nov. Previously, all authors considered this subspecies to be within the range of variability of N. b. bidentatum ( Roewer 1917, 1923; Hadži 1927, 1928, 1931; Gruber & Martens 1968; Marcellino 1973; Martens 1978; Novak et al. 1984, 1995, 1996, 2002, 2017; Novak & Gruber 2000; Komposch & Gruber 2004; Novak 2004a, 2005b, 2005c). However, Roewer (1951) noticed that Hadžiʼs individuals designated as N. bidentatum probably belonged to another species, while Novak & Gruber (2000) considered N. b. schmidti ssp. nov. a possible hybrid N. b. bidentatum × N. b. sparsum. The glans tip of a specimen from Leutschach/Lučane, (46.67° N, 15.47° E, 350 m a.s.l.; Gruber & Martens 1968: fig. 24; Martens 1978: fig. 142), Austria, is the first detailed drawing of the glans in N. b. schmidti ssp. nov. See remarks under N. pluridentatum stat. nov., N. b. gruberi ssp. nov. × N. b. schmidti ssp. nov. and N. b. martensi ssp. nov. × N. b. schmidti ssp. nov.

Distribution

Austria, Italy, Croatia, Slovenia. Vertical distribution in Slovenia: 77−2108 m a.s.l. Type locality: Dolenja Sevnica, Poljane nad Škofjo Loko (46.12° N, 14.15° E, 517 m a.s.l.), Slovenia.

Ecology

Nemastoma b. schmidti ssp. nov. is a mesophilous colline to montane subspecies occupying most regions of Slovenia, except the southern and eastern territory, inhabiting various forest and scrub communities, from mixed hardwood riverine forests in the lowland to subalpine Nardion strictae Br.-Bl. 1926 with isolated patches of Pinus mugo Turra and low-growing or krummholz communities. Phenology: adults eurychronous.

Figures

Fig. 2. The complete distributional area of the Nemastoma bidentatum Roewer, 1914 complex, according to Gruber & Martens (1968), Martens (2006), Schönhofer & Holle (2007) and own data, with marked positions of the type localities of the taxa.

Fig. 3. Distributional areas of six subspecies (circles) and hybrids (triangles, squares) within the Nemastoma bidentatum Roewer, 1914 in Slovenia and adjacent regions. Note that many localities overlap on the map.

Fig. 4. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Ch basal articles (medial, lateral views).

Fig. 5. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Pa (medial view).

Fig. 6. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Synchroscopic images of the male Ch basal article and Pa-Fe (medial views, except G, right figure, which is in dorso-medial view).

Fig. 7. Nemastoma bidentatum Roewer, 1914 complex. Penes (lateral view). Note: The penis of N. pluridentatum (Hadži, 1973) stat. nov. is unknown.

Fig. 8. Nemastoma bidentatum Roewer, 1914 complex. Glans (dorsal and lateral views). Note: The penis of N. pluridentatum (Hadži, 1973) stat. nov. is unknown.

Fig. 9. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Variation of the Ch basal article (medial view) in species, subspecies and hybrids.

Fig. 10. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Variation of the Pa-Fe (A–B first image, C, D–E first image, F–G first image; H–J, L–M right, N–O: lateral view; the others in dorso-medial view to show the armament).

Fig. 11. Nemastoma bidentatum Roewer, 1914 complex, ♀♀; Rec sem (lateral view, right portion, except F left figure, which is in dorsal view). A. The frame encloses the ovipositor portion with Rec sem. Note: Female N. pluridentatum (Hadži, 1973) stat. nov. is unknown.

Fig. 12. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. From left to right: Ch basal article (medial, lateral views), Pa (medial view) and Pe tip (dorsal, lateral views). Arrows indicate the most identifying characters. Note: Pe of N. pluridentatum (Hadži, 1973) stat. nov. is unknown.

Fig. 13. Nemastoma bidentatum Roewer, 1914 complex, ♀♀. From left to right: Silhouettes of the Ch basal article, Pa (both medial view) and Rec sem. Arrows indicate the most identifying characters. Note: female N. pluridentatum (Hadži, 1973) stat. nov. is unknown.