Nemastoma bidentatum gruberi, Novak & Novak & Kozel & Schaider & Komposch & Lipovšek & Podlesnik & Paušič & Raspotnig, 2021

Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor & Raspotnig, Günther, 2021, Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence, European Journal of Taxonomy 777, pp. 1-67 : 44

Nemastoma bidentatum gruberi ssp. nov. × N. bidentatum schmidti ssp. nov.

Figs 3 , 9M , 10M

Nemastoma bidentatum bidentatum – Gruber & Martens 1968: 141 [partim: Dugo selo]. — Martens 1978: 107 [partim: Županja, Mt. Snežnik].

Nemastoma bidentatum bidentatum × Nemastoma bidentatum sparsum – Novak & Gruber 2000: 285 [partim: Mt. Snežnik]. — Novak 2004b: 246 [partim: Dugo selo].

Diagnosis

In typical male hybrids, Ch-Apo is boomerang-shaped, Pa-Fe club-shaped, but relatively thin, with four simple, equidistant ventral Pa-Fe spines in the frontal half, Pa-Ti without proximal dorso-medial hump.

Material examined

CROATIA – WL97 • 1 ♂, 1 ♀; Zagreb − Dugo selo; J. Martens et al. leg. ( JM 837 ); PMSL • 16 ♂♂, 13 ♀♀; ibid.; 9 Sep. 2009; A. Schönhofer leg. (ASc 342, 4/2012, ASc det., TN rev. 2018); PMSL. – XL06 • 3 ♂♂, 2 ♀♀; Prevlaka; (826/2002, rev. 2018); PMSL • 4 ♂♂, 6 ♀♀; ibid.; Oct.1984; (236/2002, rev. 2018); PMSL • 2 ♂♂, 2 ♀♀; ibid.; (278/2002, rev. 2018); PMSL • 1 ♂, 1 ♀; ibid.; (280/2002, rev. 2018) • 1 ♂, 1 ♀; ibid.; 15 Aug. 1985; ( LSN 28 /1996, TN rev. 2018); PMSL • 1 ♀; ibid.; (891/2002, rev. 2018); PMSL • 1 ♂; ibid.; 30 Aug. 1985; (609/2002, rev. 2019); PMSL • 2 ♂♂; ibid.; (861/2002, rev. 2018); PMSL • 2 ♂♂, 1 ♀; ibid.; Oct.1985; (896/2002); PMSL • 1 ♀; ibid.; 7 Apr. 1986; I. Kos leg. (328/2002, rev. 2018); PMSL • 1 ♂; ibid.; (843/2002, rev. 2018); PMSL .

SLOVENIA – VL13 • 3 ♂♂, 3 ♀♀; Rakitovec ; 21 Sep. 2011; L. Slana Novak and T. Novak leg. (263/2011, rev. 2017); PMSL. – VL26 • 5 ♂♂, 2 ♀♀; Fameljska loza, Senožeče ; 1 Oct. 2011; L. Slana Novak and T. Novak leg. (350/2011, rev. 2017); PMSL • 1 ♂; Senožeče – Vremščica ; 1 Oct. 2011; L. Slana Novak and T. Novak leg. (355/2011, rev. 2017); PMSL. – VL27 • 3 ♂♂, 1 ♀; Mt. Nanos ; 1 Sep. 2011; T. Novak leg. (198/2011, rev. 2018); PMSL. – VL34 • 1 ♂, 3 ♀♀; Harije ; 19 Sep. 2011; L. Slana Novak and T. Novak leg. (224/2011, rev. 2015); PMSL • 1 ♀; Petelinje ; 27 Apr. 1994; S. Brelih leg. (815/1998, rev. 2015); PMSL. – VL58 • 2 ♂♂; Brezovica pri Borovnici ; 12 May 1997; S. Brelih leg. (83/2001, rev. 2015); PMSL. – VL67 • 1 ♂, 2 ♀♀; Lužarji ; 25 Aug. 2011; T. Novak leg. (168/2011, rev. 2018); PMSL. – VL78 • 1 ♀; Mt. Limberk ; Jun.1998; I. Bertoncelj leg. (1544/1998, rev. 2009); PMSL • 1 ♂; Veliko Mlačevo ; 13 Mar. 1992; S. Brelih leg. (813/1998, rev. 2018); PMSL. – VL79 • 1 ♂; Veliki Lipoglav ; 8 Jul. 1986; L. Slana Novak and T. Novak leg. (472/2002, rev. 2018); PMSL. – VL84 • 1 ♂, 2 ♀♀; Pleš, Borovec pri Kočevski Reki ; 19 Sep. 2013; P. Kozel and T. Novak leg. (27/2013, rev. 2015); PMSL. – VL88 • 1 ♂, 1 ♀; Krka, Krška vas pri Muljavi ; 7 Jul. 1986; L. Slana Novak and T. Novak leg. (474/2002, rev. 2018); PMSL • 1 ♀; ibid.; Jun. 1998; V. Kovačič leg. (1444/1998, rev. 2017); PMSL. – VL93 • 1 ♂; Slavski Laz ; 29 Apr. 2001; S. Brelih leg. (79/2001, rev. 2018); PMSL. – VL99 • 5 ♂♂, 1 ♀; Cirnik ; 27 May 1984; L. Slana Novak, M. Slana Novak and T. Novak leg. ( LSN 249 /1984, TN rev. 2015); PMSL • 1 ♀; Temenica ; 4 Mar. 1994; S. Brelih leg. (680a/1998, rev. 2018); PMSL. – WL04 • 5 ♂♂; Brezovica pri Predgradu ; 17 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (147/1997, rev. 2018); PMSL. – WL05 • 5 ♂♂; Črmošnjice ; 18 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (152/1997, rev. 2018); PMSL. – WL06 • 1 ♂, 1 ♀; Baza 20, Kočevske Poljane ; 17 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. ( LSN 76 /1986, TN rev. 2018); PMSL • 2 ♂♂; Podstene, Kočevski Rog ; 19 Jun. 1986 (672/2002, rev. 2017); PMSL. – WL08 • 1 ♀; Velika jama nad Trebnjem cave, Cad . No. 104, Mokronog ; 28 Jul. 2009; M. Zagmajster leg. ( PK and TN 6 /2020); PMSL. – WL09 • 1 ♂; Šentrupert; 28 Feb. 1994; S. Brelih leg. (825/1998, rev. 2018); PMSL. – WL19 • 3 ♂♂; Puščava, Mokronog ; 28 Feb. 1994; S. Brelih leg. (801/1998, rev. 2018); PMSL. – VL28 • 1 ♂; Mt. Homski hrib, Sela pri Zburah ; Jun. 1997; D. Vrček leg. (1709/1998); PMSL. – WL29 • 1 ♂, 2 ♀♀; Dolenji Boštanj ; 15 Aug. 2007; L. Slana Novak and T. Novak leg. (73/2007, rev. 2015); PMSL. – WM01 • 1 ♂, 6 ♀♀; Mt. Javor, Znojile ; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1194/2002, rev. 2018); PMSL. – WM40 • 1 ♂; Lesično ; 13 Jul. 2002; L. Slana Novak and T. Novak leg. (1130/2002, rev. 2018); PMSL • 5 ♂♂; ibid.; (1132/2002, rev. 2018); PMSL. – WM42 • 1 ♂; Mestinje ; 4 Aug. 1983; T. Novak, M. Štangelj and L. Slana Novak leg. ( LSN 44 /1983, TN rev. 2015); PMSL .

Figures

Fig. 3. Distributional areas of six subspecies (circles) and hybrids (triangles, squares) within the Nemastoma bidentatum Roewer, 1914 in Slovenia and adjacent regions. Note that many localities overlap on the map.

Fig. 9. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Variation of the Ch basal article (medial view) in species, subspecies and hybrids.

Fig. 10. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Variation of the Pa-Fe (A–B first image, C, D–E first image, F–G first image; H–J, L–M right, N–O: lateral view; the others in dorso-medial view to show the armament).