Rhaucus serripes ( Simon, 1879 )

García, Andrés F. & Kury, Adriano B., 2017, Taxonomic revision of the Andean harvestman genus Rhaucus Simon, 1879 (Arachnida, Opiliones, Cosmetidae), Zootaxa 4338 (3), pp. 401-440 : 417-422

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https://doi.org/ 10.11646/zootaxa.4338.3.1

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Rhaucus serripes ( Simon, 1879 )


Rhaucus serripes ( Simon, 1879) View in CoL comb. rest.

( Figs 1f View FIGURE 1 , 9–11 View FIGURE 9 View FIGURE 10 View FIGURE 11 , 18c View FIGURE 18 , 23 View FIGURE 23 )

Erginus serripes Simon, 1879: 204 .

Flirtea serripes: Roewer, 1912a: 77 View in CoL ; 1923: 348, figs 396–397; 1927: 593; Mello-Leitão, 1932: 78; Kury, 2003: 62.

Rhaucus (Erginus) serripes: Henriksen in Sørensen, 1932: 352 View in CoL .

Metarhaucus fuscus Pickard-Cambridge, 1905: 572 View in CoL ; Roewer, 1923: 342; 1927: 588; Kury, 2003: 70. New synonymy

Erginus fuscus: Roewer, 1912a: 68 .

Metarhaucus reticulatus Roewer, 1912b: 145 View in CoL , pl. 7, fig. 4; 1923: 343, fig. 387; 1927: 588; 1959: 80; Forcart, 1961: 51; Flórez & Sánchez, 1995: 368; Kury, 2003: 70. New synonymy

Erginus reticulatus: Roewer, 1912a: 68 .

Rhaucus (Rhaucus) geographicus Sørensen, 1932: 369 View in CoL . New synonymy

Flirtea geographica: Mello-Leitão, 1933: 110 View in CoL [by implication]; Kury, 2003: 61. Flirtea paucimaculata Roewer, 1963: 58 View in CoL , fig. 24; Flórez & Sánchez, 1995: 368; Kury, 2003: 62. New synonymy Type material. Holotype ♂ of Erginus serripes : MNHN, dry pinned, examined by photograph. Holotype ♂ of Metarhaucus fuscus View in CoL : BMNH, examined by photograph. Types of Metarhaucus reticulatus View in CoL : MHNN; SMF RI 299; ♂♀ syntypes, ERPC; 3♀ “paratypes” NHMB 63a, examined by photograph. Types of Rhaucus geographicus View in CoL : 1♂ 3♀ syntypes, ZMUC, BMNH, examined by photograph. Types of Flirtea paucimaculata View in CoL : holotype ♂, examined by photograph, 4♂ 3♀ paratypes SMF 12710; 1♂ 1♀ paratypes SMF 12716.

Distribution and habitat. COLOMBIA, Boyacá Department: Arcabuco, Villa de Leyva. Cundinamarca Department: Bogotá, Cogua, La Calera, Sesquilé, Soacha, Subachoque, Villapinzón, Zipaquirá. Between 2630 and 3400 m a.s.l., especially in Northern Andean páramo and Cordillera Oriental montane forests ecoregions ( Figs 23 View FIGURE 23 , 24a–b View FIGURE 24 ).

Diagnosis. Differs from the other species of Rhaucus by having DS with island-blots pattern ( Figs 9a View FIGURE 9 , 10a View FIGURE 10 ); DS profusely covered with granules, including tubercles of area III ( Fig. 10a View FIGURE 10 ), contrasting with smoother tubercles in all other species of Rhaucus .

Redescription. Male (ICN-AO-1376).

Measurements: CL=2, CW=4.2, AL=4, AW=5.2, BaCh=0.5, FeL III=6, FeL IV=8, TiL III=4.2, TiL IV=4.6. Intraespecific variation of body and appendages measurements in Table 1.

Dorsum ( Figs 1f View FIGURE 1 , 9 View FIGURE 9 , 10a–b View FIGURE 10 , 18c View FIGURE 18 ): Dorsal scutum alpha type, very granulate ( Figs 9a View FIGURE 9 , 10a View FIGURE 10 ), with blots that resemble islands and reach the sides of ocularium ( Figs 9a View FIGURE 9 , 10a View FIGURE 10 , 18c View FIGURE 18 ). Abdominal scutum widest at level of groove II; areas I–IV granulate. Carapace with some granules on the anterior region ( Fig. 9a View FIGURE 9 ). Ocularium low without median depression, armed with a cluster of granules ( Figs 9c–e View FIGURE 9 , 10a–b View FIGURE 10 ). Lateral margins of DS with a few granules on the medial region ( Fig. 10a View FIGURE 10 ). Mesotergum delimited, divided into four areas: area I with a pair of medium-sized paramedian tubercles; area II with a pair of small paramedian tubercles; area III with a pair of paramedian, domeshaped, coarsely granulated tubercles and another pair of medium-sized tubercles, laterally displaced; area IV with a pair of small paramedian tubercles ( Figs 9c View FIGURE 9 , 10a–b View FIGURE 10 ). Posterior border of scutum sub-straight and with a row of tubercles. Free tergites I–III with a row of tubercles ( Figs 9a, 9d–e View FIGURE 9 , 10a–b View FIGURE 10 ).

Venter ( Fig. 9b View FIGURE 9 ): Stigmatic area with a few granules. Stigmata large, oval and transverse. Coxa I with two parallel rows of tubercles increasing size distally and with an anterior smooth space for the pedipalps to rest; coxa II longer than coxa I, granulate; coxa III longer than I and II, granulate, with posterior margin sigmoid; coxa IV strongly backward, granulate. Genital operculum slightly granulate.

Pedipalps ( Figs 11a–c View FIGURE 11 ): Coxa dorsally and ventrally with one meso-distal granule. Trochanter with one dorsal short tubercle and two ventral tubercles fused at base (mesal taller); femur compressed, dorsally with a row of aligned setiferous tubercles of different sizes –not reaching the distal portion–, ventrally the same, tubercles just more separated between them. Patella distally depressed with dorsal granules; tibia depressed, spoon-shaped, dorsally granulate, with lateral borders tuberculate and some spiniferous tubercles at distal portion; inner surface slightly striated ( Fig. 11b View FIGURE 11 ). Tarsus long, conical, with some dorsal granules and ventrally with two rows of thickened setae ( Fig. 11c View FIGURE 11 ).

Chelicera ( Figs 9 View FIGURE 9 , 11d–f View FIGURE 11 ): Chelicera swollen. Basichelicerite rectangular, with a few granules, the proximal border with a row of separated tubercles and the anterior border smooth, the ectal face with two large proximal joined tubercles and a group of four distal tubercles, the dorso-distal and the basal-distal ones larger than the others, the mesal face with some small granules ( Figs 11d–e View FIGURE 11 ). Hand with one tubercle near the joint of the movable finger. Fixed finger with the inner surface finely grooved. Movable finger with one conical, sub-basal tooth and with the distal portion of the inner surface dentate ( Fig. 11f View FIGURE 11 ).

Legs ( Figs 9a–b View FIGURE 9 , 10c–e View FIGURE 10 ): Coxae I–II dorsally with one posterior process; coxa III with one proximal retrolateral tubercle; coxa IV coarsely granulate, with a prominent anterolateral clavi inguines and with a posterolateral multicapitate apophysis ( Fig. 9a View FIGURE 9 ). Trochanter I–II dorsally with some minute granules and ventrally with some tubercles; trochanter III with some dorsal and ventral granules ( Fig. 10d View FIGURE 10 ); trochanter IV more granulate than III, with one proximal tubercle and a row of ventral tubercles in retrolateral view ( Fig. 10e View FIGURE 10 ). Femora I–IV sub-straight, very granular; III–IV dorsally granulate (IV more granular than III) and ventrally with two longitudinal rows of triangular tubercles all along its length and one prolateral and one retrolateral row of tubercles, increasing in size distally: in femur III the last one tubercle thinner and in femur IV, the two last tubercles close together ( Figs 10d– e View FIGURE 10 ). Patellae I–IV dorsally very granulate, curved; patellae III–IV with the distal border coarsely granular, ventrally densely granulate and with two large tubercles on the retrolateral-ventral side. Tibiae I–IV granulate; tibiae III–IV strongly curved, with two longitudinal rows of tubercles –of different sizes– very close together in all the length (in tibia IV the distal tubercle is fused and the prolateral row of tubercles almost joins the ventral row) ( Figs 10d–e View FIGURE 10 ). Metatarsus I–IV with one ventrodistal seta. Tarsi I–II with one smooth claw; tarsi III–IV with 2 subparallel smooth claws and a tarsal process ( Fig. 10c View FIGURE 10 ). Tarsal counts: 6(3)–6(3)/13(3)–15(3)/9–9/11–11. Intraspecific variations in Table 2.

Color (in ethanol): Carapace Strong Reddish Brown (40), reticulated on the ocularium. Areas I–IV Dark Reddish Brown (44). Blots in carapace Brilliant Greenish Yellow (98). Pedipalps, coxae I–IV, free tergites and chelicera Strong Reddish Brow n (40). Trochanters I–IV and legs I–III Strong Yellowish Brown (74). Legs IV Strong Reddish Brown (40).

Male genitalia ( Figs 11f–h View FIGURE 11 ): VP of penis subrectangular with very concave distal border; VP with two lateral, elongated, dense patches of type 4 microsetae and a longitudinal gap in between ( Fig. 11h View FIGURE 11 ). VP with two apical MS C curved and laterally inserted; two MS D, the most distal large and straight, following the same lateral row as MS C; the other (D2) smaller, inserted dorso-laterally on the middle third of VP; two MS A straight and laterally inserted, near D2 ( Fig. 11g View FIGURE 11 ); two small MS E volcano-shaped on the ventral face, close together; one small MS B volcano-shaped inserted laterally at the base of the VP ( Fig. 11h View FIGURE 11 ). Glans mostly smooth; stylus long, tip surface tuft-shaped, fin-like process ( Fig. 11f View FIGURE 11 ).

Female. Similar to male, but differs by having carapace at areas I–IV level more globose; anterior part of carapace narrower and coda divergent; tubercles of area III taller; legs III–IV thinner; tubercles in ventral rows of legs III–IV smaller; chelicerae are not hyperthelic in females ( Figs 9d–e View FIGURE 9 ).

Other material examined. 60 specimens (31 males, 28 females, 1 juv.) in 25 lots. COLOMBIA: Boyacá Department: Villa de Leyva , 1♂ 3♀ (ICN-AO-262), S.F.F. Iguaque sector Carrizal [05°41'N 73°27'W], 2800 m, 31.xii.2004, E. Flórez, G. Giribet & M. Porras leg. GoogleMaps ; 3♂ 2♀ (ICN-AO-298), S.F.F. Iguaque sector Carrizal [05°41'N 73°27'W], 2900 m, 9.vi.2001, E. Flórez leg. GoogleMaps ; 3♂ 1♀ (ICN-AO-1376), S.F.F. Iguaque [5.720016N - 73,457901W], 2800 m, 22.ix.2013, A. García & S. Galvis leg. GoogleMaps ; 1♂ (IAvH-3000018/IAvH-I-4), S.F.F. Iguaque , lagoon shore [05°38'N 73°29'W], 3450 m, 13.ix.1998, E. González leg. GoogleMaps ; 1♀ (IAvH-3000073/IAvH-I-3), S.F.F. Iguaque, Cabaña Carrizal [05°25'N 73°27'W], 2850 m, 22–24.iv.2001, P. Reina leg. GoogleMaps ; 1♂ (IAvH-3000006/IAvH-I-11), S.F.F. Iguaque , lagoon shore [05°38'N 73°29'W], 3340 m, 14.x.1998, S. Sierra leg. GoogleMaps ; 4♂ 2♀ (MNRJ 19377), S.F.F. Iguaque [05°42'N 73°27'W], 2941 m, A. García & A.B. Kury leg.; Arcabuco, 1♀ (IAvH-3000020/IAvH-I-105), close to El Cometa [05°44'N 73°29'W], 3000 m, iv.1998, D GoogleMaps . Dávila & E. González leg. Cundinamarca Department: La Calera , 1♀ (ICN-AO-85) [4°43'N 73°58'W], 2900 m, 25.v.1978 GoogleMaps ; 1♂ 1♀ (ICN-AO-94), Bosque El Granizo , 3200 m, 3.iii.1991, O. Muñoz & K. Estupiñan leg. ; 1♀ (ICN-AO-480), Finca La Esperanza [4°42'N 73°57'W], 29.x.2005, M. Romero leg. GoogleMaps ; 1♂ (ICN-AO-1009), vía a Mundo Nuevo [4°38'4N 73°54'W], 2850 m, 17.x.2000, E. Flórez leg. ; 1♂ (ICN-AO-293), P.N.N. Chingaza , 3400 m, 20.vii.1987 ; 1♀ (ICN-AO-426), P.N.N. Chingaza, Valle de los Frailejones [04°62'N 73°72'W], 24.iii.2003, J. Cepeda leg.; Zipaquirá , 2♀ (ICN-AO-215), Vereda Ventalarga, Represa Pantanoredondo [5°2'N 74°2'W], 3000 m, 27.iv.2003, E. Flórez & UN-Octapodos group leg.; Sesquilé, 1♀ (ICN-AO-239), Cerro Las Tres Viejas [5°1'N 73°47'W], 3250 m, 28.i.2001, J. Betancur leg.; Subachoque GoogleMaps , 1♂ (ICN-AO-400), Vereda El Tobal, Finca El Cerro [4°55'N 74°12'W], 2900 m, 28.vii.2002, F GoogleMaps . Gomez leg.; Villapinzón, 1♂ (ICN-AO-440), Páramo de Guacheneque, camino al Pozo del Oso [05°10'N 73°33'W], 7.iv.2012; Sibaté GoogleMaps , 1♀ (ICN-AO-486), San Miguel, antes del Alto del Cuchuco [4°26'N 74°17'W], 2630 m, 1.xii.1979, UN biology students leg.; Bogotá GoogleMaps , 1♂ 1♀ (ICN-AO-1039), vía a la Vírgen de la Peña [4°34'N 74°3'W], 2700 m, 12.v.2012, M. Medrano & A. Garcia leg. GoogleMaps ; 2♂ 3♀ (ICN-AO-995), Club Campestre , 22.vi.1997, F . López leg.; 1♂ 1♀ ( MNRJ 8711 View Materials ), Páramo de Cruz Verde , 4.x.1992, D . Vera leg.; 1♂ 1♀ (MNRJ 00175)[ Meterginus uniformis M-L in schedula]; Cogua, 2♂ 3♀ (ICN-AO-1212), Parque Forestal Embalse del Neusa [5.141987, -73.962285], 3100 m, 3.xi.2013, C. Perafán & D. Martínez leg.; Soacha , 1♂ (ICN-AO-1440), Bogotá-Silvania road, near Chusacá [4°32'N 74°16'W], 1.xii.1979, G group leg. GoogleMaps














Rhaucus serripes ( Simon, 1879 )

García, Andrés F. & Kury, Adriano B. 2017

Flirtea geographica: Mello-Leitão, 1933 : 110

Kury 2003: 61
Kury 2003: 62
Florez 1995: 368
Roewer 1963: 58
Mello-Leitao 1933: 110

Flirtea serripes:

Kury 2003: 62
Mello-Leitao 1932: 78
Roewer 1912: 77

Erginus fuscus:

Roewer 1912: 68

Metarhaucus reticulatus

Kury 2003: 70
Florez 1995: 368
Forcart 1961: 51
Roewer 1912: 145

Erginus reticulatus:

Roewer 1912: 68

Metarhaucus fuscus

Kury 2003: 70
Roewer 1923: 342
Pickard-Cambridge 1905: 572

Erginus serripes

Simon 1879: 204
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