Zagryphus tilloyi (Theobald, 1937) Spasojevic & Broad & Klopfstein, 2022
publication ID |
https://dx.doi.org/10.3897/fr.25.83034 |
publication LSID |
lsid:zoobank.org:pub:6402F8F1-5229-4153-823F-CAEA106F90A1 |
persistent identifier |
https://treatment.plazi.org/id/54ACF8C1-C3AA-594B-B8D6-528113AC92FC |
treatment provided by |
by Pensoft |
scientific name |
Zagryphus tilloyi (Theobald, 1937) |
status |
comb. nov. |
Zagryphus tilloyi (Theobald, 1937) comb. nov.
Fig. 17 View Figure 17
* Promethes tilloyi Theobald, 1937
Material.
Holotype (part only, #Am_8 / MNHN.F. B24392 View Materials ) examined at MNHN. F. Photograph obtained from MNHN.F.
Stratum.
Aix-en-Provence, Bouche-du-Rhône, France. Late Oligocene (Chattian), 28.4-23.0 Ma.
Description.
Female. Holotype in lateral to ventrolateral view with nearly complete antennae, head, mesosoma with details indistinct; fore wings darkened and on top of each other, venation in part difficult to discern, one hind wing; at least parts of all legs, and metasoma with ovipositor with sheaths. Body length 6.1 mm.
Dark brown, antennae reddish at base and with white bands around mid-length; with reddish colouration on head and mesosoma, wings strongly tinted, brown; legs dark brown; metasoma dark, but possibly with irregular lighter markings on T2, ovipositor orange, its sheaths dark brown.
Head quite round, with large eyes, with a nose-like protrusion where clypeus would be. Antennae 4.5 mm, with at least 28 (probably around 32) flagellomeres, basal ones elongate, those apically of white band quadrate to transverse. Mesosoma not well preserved, with mesosternal scrobe, indication of epicnemial carina and notauli reaching at least over first third of mesoscutum. Metapleuron as long as wide, with submetapleural carina complete; propodeum with pleural carina, lateral longitudinal and maybe other carinae complete, meeting point of lateral longitudinal and posterior transverse carina might be thickened (cf. propodeal apophyses). Fore wing 3.6 mm, rather stout; areolet open, 2Rs rather short; 1cu-a meeting M + Cu opposite of 1M; 1M & 1Rs bowed inwards; 3Cu more than 3 × longer than 2cu-a, strongly inclivous; cell 2R1 2.9 × longer than wide. Hind wing with 1Rs about as long as rs-m and 1Cu much longer than cu-a. Legs slender, hind femur 4.4 ×, hind tibia 7 × longer than wide, dark dorsally light ventrally. Metasoma compressed from T4 or T5; T1 not well preserved, but slightly elongate and tapering towards base, following tergites transverse; sternites quite well sclerotized; hypopygium triangular in lateral view. Ovipositor 1.9 mm, about 1.3 × as long as hind tibia, curved downwards; ovipositor sheaths widened around mid-length.
Interpretation.
Even though many characters are not clearly visible, the medially expanded ovipositor sheaths firmly place this fossil in the Tryphoninae tribe Oedemopsini . Indeed, the specimen shows a remarkable similarity to the extant Zagryphus nasutus (Cresson, 1868): the nose-like expansion of the clypeus, inclivous 3Cu in the fore wing and short cu-a in the hind wing, triangular but not strongly elongate hypopygium, and down-curved ovipositor with medially expanded sheaths support a placement in this genus. Even the colouration is a good match, including the white band on the antenna and orange base, a pattern often observed in Zagryphus and related genera. Zagryphus tilloyi can be distinguished from the other members of the genus by its smaller size (at least 4.8 mm in extant species) and more strongly tinted wings.
Xoridinae Shuckard, 1840
Xorides Latreille, 1809
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
Kingdom |
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Phylum |
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Class |
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Order |
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Family |
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SubFamily |
Acaenitinae |
Genus |
Zagryphus tilloyi (Theobald, 1937)
Spasojevic, Tamara, Broad, Gavin R. & Klopfstein, Seraina 2022 |
* Promethes tilloyi
Spasojevic & Broad & Klopfstein 2022 |