Thaumastoptera (Thaumastoptera) intermixta Savchenko, 1974

Starý, Jaroslav & Oboňa, Jozef, 2018, Palaearctic species of Thaumastoptera (s. str.) Mik (Diptera: Limoniidae), Zootaxa 4394 (2), pp. 227-234 : 230-231

Thaumastoptera (Thaumastoptera) intermixta Savchenko, 1974

(Figs. 2, 7, 8)

Thaumastoptera intermixta Savchenko 1974: 34 (description), Fig. 34 (male terminalia).

Diagnosis. Small species, yet slightly larger than other species treated here. Body colouration in general pale yellow. Wing without any darker marks. Venation with upper branch of medial fork about 8 times as long as its petiole. Femora and tibiae without any darker rings. Male terminalia with aedeagus exceedingly long, by about one third its length longer than gonocoxite, tapered into subacute tip ( Figs. 7, 8 ). Wing length 5.2–5.6 mm.

Material examined: 2 males, 1 female.

Russia: Krasnodar Territory, Sochi / Khosta (43°31'24"N 39°52'47"E), 15.v.2011, male (D. Gavryushin leg.) ( JSO) . Georgia: “Autonomous Soviet Socialist Republic of Adjara: …, …, 16.vi.1978, E. Savchenko leg.” [handwritten label in Cyrilic, illegible in part], male, female ( JSO) .

Redescription. Male: Head. Suffused with dark greyish brown pruinosity on frons and vertex. Rostrum and palpus brown. Antenna 16-segmented, moderately long, reaching to about base of wing, brown throughout. Flagellomeres spherical proximally, passing into ovoid and narrower distally, with verticils moderately dense, longest ones about twice as long as their respective flagellomeres.

Thorax. Generally yellow, mostly shiny dorsally, including pronotum, scutellum, and mediotergite. Pleuron likewise shiny yellow. Wing membrane tinged yellowish, veins yellow. No dark spots or darkly tinged sections of veins on wing. Venation generally usual for Thaumastoptera s. str. Sc1 ending about opposite slightly more than one third length of Rs; upper branch of medial fork (M1+2) differing from other species in being very long, about 8 times as long as its petiole (M1+2+3), or longer (Fig. 2). Halter yellow throughout. Legs pale yellow, without any darker rings.

Abdomen. Yellow. Male terminalia ( Figs. 7, 8 ) yellow. Segment 9 (basal ring) simple, more or less straight at dorsal posterior margin. Proctiger generally semioval, in part covered with spinulae, broadly rounded at posterior margin. Gonocoxite moderately long, slightly arched. Gonostylus rather long, produced into slightly curved, setose beak, directed medially and subacute at apex, and with another, broadly obtuse projection on outer margin of gonostylus before apex; latter projection with a few spinoid setulae. Group of longer setae at inner margin, at about half-length of gonostylus. Aedeagus exceedingly long, by about one third its length longer than gonocoxite, generally parallel-sided, rather broad proximally, abruptly narrowed to about half its breadth at its half-length, apex of aedeagus tapered into subacute tip. Paramere very short, just as lobe at base of aedeagus.

Female. Resembling male in general appearance.

Distribution. Russia: North Caucasus; Georgia. Distribution in Uzbekistan and Tajikistan given by Oosterbroek (2017) probably is not based on published primary data (V. Lantsov, pers. comm.).

Discussion. Externally, T. (T.) intermixta differs from the other species treated here by its slightly larger size and its completely yellow body colouration, without any dark marks on the wings and any apical rings on the femora and tibiae. Male terminalia are conspicuous in having a very long aedeagus ( Figs. 7, 8 ).

Figures

FIGURES 5–8. Male terminalia.5–6: T.(T.) calceata (Czech Republic:Moravia), general view (5) and aedeagal complex (6), dorsal; 7–8: T. (T.) intermixta (Georgia: Adjara), general view (7) and aedeagal complex (8), dorsal. Scale bars 0.25 mm. Abbreviations: gc, gonocoxite; gs, gonostylus; ptg, proctiger; s9, segment 9.