Peckia (Pattonella) kladosoides, Buenaventura & Pape, 2013

Buenaventura, Eliana & Pape, Thomas, 2013, <strong> Revision of the New World genus <em> Peckia </ em> Robineau-Desvoidy (Diptera: Sarcophagidae) </ strong>, Zootaxa 3622 (1), pp. 1-87 : 40-41

Peckia (Pattonella) kladosoides n. sp.

( Figs. 15, 16 , 58 )

Description. Male. Length. 14.3 mm. Head. Ocellar setae equal to or smaller than postoculars. Outer vertical seta of same size as postoculars. Orbital setae black. Four frontal setae situated below the dorsal limit of the lunule in lateral view. Genal setae black anteriorly, yellow or white posteriorly. First two rows of occipital setae black, others yellow. Thorax. Chaetotaxy: acrostichals = 0 + 1, dorsocentrals = 0 + 2 (anterior one shorter), intra-alars = 2 + 2 (anterior one shorter), supra-alars = 2 + 3, basal scutellars = 4. Prosternum and posterior surface of hind coxa with yellow or light brown setae. Antero-ventral scutellar setae black. Three katepisternal setae. Postalar wall with white setae anteriorly. Lower calypter without dorsal setae, with a central dark spot and a fringe of long hair-like setae along outer margin, extending to its posterior margin. Mid femur with a ctenidium. Antero-dorsal surface of mid tibia with 2 median setae and 1 apical seta. One row of setae on ventral surface of hind femur. Hind tibia with 2 setae in the basal third, 1 in the middle third and 1 preapical on antero-dorsal surface. Abdomen. Postero-ventral seams between T3/T4 curved anteriorly, so that segments are markedly longer dorsally than ventrally. Posterior seam of T5 projected posteriorly and ventrally. Postero-lateral area of T5 with setae directed towards the ventral body region. T5 with golden microtrichosity. Abdomen in lateral view with golden microtrichosity. ST1+3 with yellow and white setae. Two lateral setae on each side of T4. Terminalia. ST5 brown. Medial margin of ST5 Λ- shaped. Inner margins of ST5 arms straight. ST5 arms with rounded apophyses anteriorly. Inner margins of ST5 arms with a patch of short setae on the median region. Syntergosternite 7+8 as long as high in lateral view, orange, with golden microtrichosity. Epandrium bright orange. Cercus orange proximally and brown or dark brown distally. Dorsal margin of cercus strongly angulated at about midlength, and an acute apex in lateral view. Surstylus black, finger-shaped. Pregonite triangular, becoming narrower towards the apex, with an acute apex and straight in lateral view. Postgonite elongated, with a hooked apex. Basi- and distiphallus connected by a desclerotized strip. Phallic tube curved and extremely elongated. Juxta with a demarcated connection with the phallic tube. Juxta without juxtal lateral plate, semi-sclerotized, composed of two lobes both compressed dorso-ventrally. Distiphallus with a pair of lateral styli separated, each with a longitudinal cleft, not visible in lateral view. Vesica strongly developed, composed of two slender filaments with an angled section at mid length, their length exceeding the length of the juxta.

Female. Unknown.

Variable features. The following differences were observed in the paratypes as compared with the holotype: some specimens present asymmetry in the number of setae situated below the dorsal limit of the lunule, with 4 on the right side and 5 on the left.

Taxonomic remarks. Peckia (Pattonella) intermutans has a broad, trapezoidal pregonite, while this is more slender and more triangular in P. (Pattonella) kladosoides . Peckia (Pattonella) kladosoides differs from P. (Pattonella) resona by the absence of hair-like setae on the dorsal surface of lower calypter. The juxta in P. (Pattonella) kladosoides is smaller and more flattened than in P. (Pattonella) smarti , P. (Pattonella) intermutans , and P. (Pattonella) resona , but its vesica is more elongated.

Biology. The type specimens were collected on decomposed fish in a tropical wet secondary forest between 843 and 1285 m of altitude, with a 24° C average temperature and an annual rainfall of 4740 mm.

Distribution. NEOTROPICAL— Colombia (Boyacá).

Etymology. The specific epithet is a feminine adjective deriving from the Greek word klados, meaning branch, twig or stem. The specific epithet refers to the vesica, which resembles the branch of a tree.

Holotype. Male, COLOMBIA, Boyacá, Santa María, La Almenara , Bosque , decomposed fish, 4°53'0.2''N 73°15'15''W, 1285m, 27.xii.2008, E. Buenaventura ( ICN).

Paratypes. 1 male, same data as holotype ( MNRJ) ; 1 male, COLOMBIA, Boyacá, Santa María, Icacuye, Bosque , decomposed fish, 4°53'42.8''N 73°16'43.1''W, 843m, 28.xii.2008, E. Buenaventura ( ZMUC) .

Figures

FIGURES 8–16. 8–9. Peckia (Peckia) hirsuta. 8. Lateral styli and possible remnant of median stylus, apical view. 9. Reduced median stylus. 10. Peckia (Pattonella) intermutans. Lateral styli and median membranous opening, apical view. 11–12. Peckia (Squamatodes) ingens. 11. Lateral styli fused, apical view. 12. Lateral styli fused with an arrow pointing to the median seam, dorsal view. 13. Peckia (Peckia) gulo. Left lower calypter, dorsal view. 14. Peckia (Euboettcheria) anguilla. Distiphallus, left lateral view. 15–16. Peckia (Pattonella) kladosoides n. sp. 15. Distiphallus, left lateral view. 16. Lateral styli, left lateral view. (Abbreviations: J—juxta, Ls—lateral stylus, Mo—median opening, rMS—remnant of median stylus, V—vesica.)

FIGURES 54–65. 54. Peckia (Euboettcheria) buethni. Male terminalia with an arrow pointing to the fin-shaped crest, left lateral view. 55. Peckia (Pattonella) intermutans. Male terminalia with the lower arrow pointing to the small, acute projection of the distiphallus, left lateral view. 56–57. Peckia (Pattonella) resona. 56. Male terminalia, left lateral view. 57. Lower calypter, dorsal view, with an arrow pointing to the dorsal setae. 58. Peckia (Pattonella) kladosoides n. sp. Distiphallus, left lateral view. 59. Peckia (Pattonella) pallidipilosa. Male terminalia, left lateral view. 60–62. Peckia (Peckia) ecuatoriana. 60. Distiphallus, left dorso-lateral view. 61. Distiphallus, apical view. 62. Male terminalia (distiphallus missing), left lateral view. 63–64. Peckia (Peckia) sarmientoi n. sp. 63. Distiphallus, apical view. 64. Male terminalia, left lateral view. 65. Peckia (Peckia) nigricauda. Male terminalia with an arrow pointing to the hooked projection of the lateral plates, left disti-lateral view. (Abbreviations: aJ—juxtal accessory, Ce—cercus, J—juxta, Jlp—juxtal lateral plate, Ls –lateral stylus, Pog—postgonite, Prg—pregonite, Sr—surstylus, V—vesica.)