Glyptocombus Heidemann, 1906

Weirauch, Christiane, Hoey-Chamberlain, Rochelle & Knyshov, Alexander, 2018, Synopsis of Schizopteridae (Hemiptera, Heteroptera, Dipsocoromorpha) from the United States, with description of seven new species from the US and Mexico, ZooKeys 796, pp. 49-82: 49

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Glyptocombus Heidemann, 1906


Glyptocombus Heidemann, 1906  Figures 1, 2, 3, 10

Type species.

Glyptocombus saltator  Heidemann, 1906

Revised diagnosis.

Recognized among genera of New World Hypselosomatinae  by dense setation on all surfaces of the tibia, extending from base to apex, first labial segment not dorsally expanded, and third and fourth labial segments without processes.

Revised description.

Male (Figure 1): macropterous or with elytriform forewing, length: 1.14-1.74 mm; body ovoid. Coloration: general coloration tan to black. Surface and Vestiture: scape with 5 setae, clypeus with 3 long, stout setae, buccula and first labial segment each with pair of stout ventrolateral setae, second labial segment with pair of lateral setae (Figure 1), long, stout seta dorsoapically on third tarsal segment of all legs, anophore without distinctive setae (Figure 3A). Structure: Head: first labial segment not enlarged, clypeus simple. Thorax: tarsal formula 2-3-3, parempodia long and thin, bladder-like arolium absent. Abdomen: abdomen with 6 spiracles on left and 5 on right side, right side of tergum 8 with small projection containing spiracle, left side with large mitten-shaped projection with spiracle and long seta, sterna 7 and 8 distinct, posterior margin of sternum 7 simple, left side with triangular flap, sternum 8 rectangular-ovoid (Figure 3B). Genitalia (Figure 3A): right paramere long, stout with rounded apex and short, rounded process at base, left paramere scapula-shaped with large thumb-shaped projection, vesica of average length with one coil and small rounded conjunctival appendage at base, anophoric process crab claw-shaped, extending posteriorly on left side.

Female (Figure 1): with elytriform forewing, length: 1.42-1.51 mm; body ovoid. Coloration: general coloration tan to black. Surface and vestiture: scape with 5 setae, setae on clypeus, buccula and first and second labial segments as in male. Structure. Head: first labial segment and clypeus as in male. Thorax: tarsal formula 2-2-3, pretarsus as in male (Figure 1). Genitalia (Figure 3C, D): anterior gonapophysis with 3 teeth, without subapical serration, posterior gonapophysis with two teeth, median styloid bifurcate, gonoplac small, oriented ventrad, spermathecal gland spherical; spermathecal gland duct straight, spermathecal reservoir globular with one bend, spermathecal duct short and relatively straight.


This is the only genus of Hypselosomatinae  with almost exclusive Nearctic distribution based on locality records of the single previously described and three new species, as well as one female specimen from Chiapas that represents an undescribed species. Specimens have been collected using Berlese extraction, hand collecting, yellow pan traps, pitfall traps (single and array, normal and dung), sifting leaf litter, suction traps, and UV lighting. Extracted substrates included beech humus, bottomland hardwood remnant, fallen leaves, “rubbish,” earth, and the forest floor of Palmetto-gumbo limbo upland.