BARYPINA

ROIG-JUÑENT, SERGIO, 2000, The Subtribes And Genera Of The Tribe Broscini (Coleoptera: Carabidae): Cladistic Analysis, Taxonomic Treatment, And Biogeographical Considerations, Bulletin of the American Museum of Natural History 2000 (1), pp. 1-90 : 47-48

BARYPINA JEANNEL, 1941

TYPE GENUS: Barypus Dejean, 1828 . TAXA INCLUDED: Jeannel (1941) considered this group to include Barypus and the Australian and New Zealand genera of Broscini that have the basal orifice of median lobe completely closed. However, Ball (1956) included only Barypus . I considered this subtribe as Ball conceived it, to include only Barypus and a new genus described herein. The other genera considered by Jeannel are included in the subtribe Nothobroscina .

Barypina includes 23 species in two genera, Barypus and Microbarypus new genus (monospecific) both from southern South America. consider Microbarypus at generic rank, and not a subgenus of Barypus because the two are largely different. Barypus shows several synapomorphies that traditionally define its species, and the inclusion of Microbarypus would imply a complete different and new redefinition of Barypus .

Barypina is the sister group of Creobiina . In both subtribes, the internal sac has lost the sclerites X and Y and the apical plate; also, the female genital tract has lost its vaginal apophysis. In terms of synapomorphies, Barypina is among the most distinctive groups in the entire tribe, as shown in figure 15. Besides sharing several features with Creobiina and allied genera, Barypina shows outstanding similarities to the Oregus clade in the structures of the mouthparts. The basal keel on the median lobe, a character used by Ball (1956) to define the subtribe Barypina , is present at different degrees in the genera Oregus (figs. 159–160) and Percosoma (fig. 170), indicating that this structure may have evolved independently.

The barypine species live in different habitats. The species of Barypus live in grassland or shrubby arid habitats, while Microbarypus inhabits the Nothofagus forest.