Anomala consanguinea, Zhao, 2019

Zhao, Ming Zhi, 2019, A new species of the genusAnomala from southern China (Coleoptera: Scarabaeidae: Rutelinae), Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (2), pp. 481-489 : 482-487

Anomala consanguinea sp. nov.

( Figs 1A–F ; 3A–C ; 4A–C, G ; 5A–C )

Type locality. China, Hainan Province, Ledong Li Autonomous County, Jianfeng Town, Jianfengling National Forest Park, Tianchi, 950m.

Type material (608 spec.). HOLOTYPE: J ( SCAU), ‘China: Hainan, Ledong Li / Mount Jianfengling / ƾ池 [= Tianchi], 950m / 2009.IV.10 / Wen-Hsin Lin leg.’ PARATYPES: CHINA: 2 JJ ( CCTC), ‘China: Hainan, Ledong Li / Mount Jianfengling / Tianchi ƾ池 [= Tianchi], 950m / 2010. IV.21 / Wen-Hsin Lin leg.’; 1 J ( CCTC), ‘China: Hainan, Ledong Li, / Mount Jianfengling, / Observation Tower, / 1000m, 2009.IV.26, / Wen- -Hsin Lin leg.’; 2JJ 7♀♀ ( ZMGC), ‘China: Hainan, Ledong Li, / Mount Jianfengling, / Mingfenggu ńNī [= Mingfenggu] / 2015.IV.23, Lu Qiu leg.’; 3 JJ ( CCTC), ‘China: Hainan, / Mount Wuzhishan, 水满乡 [= Shuimanxiang Township] / 700m, 2010.IV.10, / Wen-Hsin Lin leg.’; 1 J 1 ♀ ( CCTC), ‘China: Hainan, / Mount Wuzhishan, / 700m, 2011. IV.21, / Yi-Ting Chung leg.’; 1J ( ZMGC), ‘China: Hainan, Qiongzhong County / Yinggeling Nature Reserve, / Yinggezui, 656m, 2011.IV.26, / Yi-Ting Chung leg.’; 1 J 1 ♀ ( ZMGC), ‘China: Hainan, / Diaoluoshan Mt., / 2015.IV.16-18, / Lu Qiu leg.’; 4 JJ 1 ♀ ( IZAS), ‘ 海南‡中l母 山 [= Hainan, Qiongzhong, Mount Limushan] / 七¶村 [= Qijiacun Village] 645m / 2010.IV.3 / 中LJnḿ物ıffñ [= IZAS] // 19.17° N / 109.72° E / ze人：hxẉ [= Mei-Ying Lin leg.] / 中LJnḿ物ıff ñ [= IZAS] // IOZ(E)1948598, IOZ(E)1948600–IOZ(E)1948603’; 15 JJ 5 ♀♀ ( IZAS), ‘ 海南‡中l母山 [= Hainan, Qiongzhong, Mount Limushan] / 七¶村 [= Qijiacun Village] 645m / 2010.IV.4 / 中LJnḿ 物ıffñ [= IZAS] // 19.17° N / 109.72° E / ze人：hxẉ [= Mei-Ying Lin leg.] / 中LJnḿ物ıffñ [= IZAS] // IOZ(E)1948073– IOZ(E)1948075, IOZ(E)1948077–IOZ(E)1948078, IOZ(E)1948080– IOZ(E)1948082, IOZ(E)1948086, IOZ(E)1948689–IOZ(E)1948690, IOZ(E)1948147–IOZ(E)1948151’; 1 J 2 ♀♀ ( IZAS), ‘ 海南‡中l母 山 [= Hainan, Qiongzhong, Mount Limushan] / 七¶村 [= Qijiacun Village] 647m / 2010.IV.6 / 中LJnḿ物ıffñ [= IZAS] // 19.17° N / 109.72° E / ze人：hxẉ [= Mei-Ying Lin leg.] / 中LJnḿ物ıff ñ [= IZAS] // IOZ(E)1948604, IOZ(E)1948607, IOZ(E)1948609’; 5 JJ 3 ♀♀ ( IZAS), ‘ 海南‡中l母山 [= Hainan, Qiongzhong, Mount Limushan] / 七¶村 [= Qijiacun Village] 657m / 2010.IV.7 / 中LJnḿ 物ıffñ [= IZAS] // 19.17° N / 109.71° E / ze人：hxẉ [= Mei-Ying Lin leg.] / 中LJnḿ物ıffñ [= IZAS] // IOZ(E)1948083, IOZ(E)1948103–IOZ(E)1948106, IOZ(E)1948111, IOZ(E)1948113– IOZ(E)1948114’; 15 JJ 16 ♀♀ ( IZAS), ‘ 海南‡中l母山 [= Hainan, Qiongzhong, Mount Limushan] / Ø河护hü [= Yinghe Forestry Preserve Station] 641m / 2010.IV.5 / 中LJnḿ物ñ [= IZAS] // 19.18° N / 109.74° E / ze人：hxẉ [= Mei-Ying Lin leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1948657–IOZ(E)1948687’; 1 J ( IZAS), ‘ 海南‡中 县l母山 [= Hainan, Qiongzhong, Mount Limushan] / 七¶村 [= Qijiacun Village] 645m / 2010.IV.3 / 中LJnḿ物ıffñ [= IZAS] // 19.17° N / 109.72° E / ze人：张ƚhv [= Kui-Yan Zhang leg.] / 中LJnḿ物 ıffñ [= IZAS] // IOZ(E)1948112’; 2 ♀♀ ( IZAS), ‘ 海南‡中l母山 [= Hainan, Qiongzhong, Mount Limushan] / 海Ɲ： 640m / 2007.V.15 李娴 [= Xian Li leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1947920, IOZ(E)1947978’; 32 JJ 5 ♀♀ ( IZAS), ‘ 海南白沙县ƶ哥岭 [= Hainan, Baisha County, Yinggeling] / 海Ɲ： 790m [= alt. 790m] / 2010.IV.13 / 中LJnḿ物ñ [= IZAS] // 18.97° N / 109.29° E / ze人：hxẉ [= Mei-Ying Lin leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1948610, IOZ(E)1948613– IOZ(E)1948616, IOZ(E)1948619–IOZ(E)1948621, IOZ(E)1948044–IOZ(E)1948068, IOZ(E)1948070–IOZ(E)1948072, IOZ(E)1948087’; 28 JJ 8 ♀♀ ( IZAS), ‘ 海南白沙县ƶ哥岭 [= Hainan, Baisha County, Yinggeling] / ƶ哥¬ [= Yinggezui] 627m / 2010.IV.17 / 中LJnḿ物ñ [= IZAS] // 19.05° N / 109.56° E / ze人：hxẉ [= Mei-Ying Lin leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1948611, IOZ(E)1948622–IOZ(E)1948656’; 14 JJ 7 ♀♀ ( IZAS), ‘ 海南省白沙 ƶ哥岭 [= Hainan, Baisha County, Yinggeling] / 2011-V-1 IJa [= lighting trap] / hĸ信 [= Wen-Hsin Lin leg.] 600m / 中国LJfnḿ物 ñ [= IZAS] // IOZ(E)1948692–IOZ(E)1948696, IOZ(E)1948698– IOZ(E)1948706, IOZ(E)1948709–IOZ(E)1948715’; 1 J ( IZAS), ‘ 海南 省白沙ƶ哥岭 [= Hainan, Baisha County, Yinggeling] / 2011-IV-27 ~30 / 600-780m hĸ信 [= Wen-Hsin Lin leg.] / 中国LJfnḿ物ñ [= IZAS] // IOZ(E)1948708’; 4 JJ 3 ♀♀ ( IZAS), ‘ 海南白沙县ƶ哥岭 [= Hainan, Baisha County,Yinggeling] / 南开乡 [= Nankaixiang Township]

336m / 2010.IV.16 / 中LJnḿ物ñ [= IZAS] // 19.01° N / 109.36° E / ze人：hxẉ [= Mei-Ying Lin leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1948108–IOZ(E)1948110, IOZ(E)1948116–IOZ(E)1948118’; 1 f * ( IZAS), ‘ 海南白沙县ƶ哥¬ [= Hainan, Baisha County, Yinggezui] / 2009.V.6 IJa [= lighting trap] / M婷玉 [= Ting-Yu Hu leg.] / 中LJ nḿ物ñ [= IZAS] // IOZ(E)1947910’; 1♀ ( IZAS), ‘ 海南白沙县ƶ哥 岭 [= Hainan, Baisha County,Yinggeling] / ƶ哥¬保护ü [= Yinggezui Station] IJa [= lighting trap] / 2009.V.6 张ł [= Rui Zhang leg.] / 中 LJnḿ物ñ [= IZAS] // IOZ(E)1948009’; 1 J ( IZAS), ‘ 海南白沙县ƶ 哥岭 [= Hainan, Baisha County, Yinggeling] / Ʋ坝水库 [= Hongba Reservoir] IJa [= lighting trap] / 2009.V.3 M婷玉 [= Ting-Yu Hu leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1947997’; 5 ♀♀ ( IZAS), ‘ 海南白 沙县ƶ哥岭 [= Hainan, Baisha County, Yinggeling] / Ʋ坝水库 [= Hongba Reservoir] IJa [= lighting trap] / 2009.V.4 M婷玉 [= Ting-Yu Hu leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1947911–IOZ(E)1947915’; 1 J 3 ♀♀ ( IZAS), ‘ 海南白沙县ƶ哥岭 [= Hainan, Baisha County, Yinggeling] / Ʋ坝水库 [= Hongba Reservoir] IJa [= lighting trap] / 2009.V.5 M婷玉 [= Ting-Yu Hu leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1947998, IOZ(E)1948000–IOZ(E)1948002’; 1 ♀ ( IZAS), ‘ 海南 白沙县ƶ哥岭 [= Hainan, Baisha County, Yinggeling] / Ʋ坝水库 [= Hongba Reservoir] IJa [= lighting trap] / 2009.V.4 张ł [= Rui Zhang leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1948008’; 1 J 2 ♀♀ ( IZAS), ‘ 海南白沙县ƶ哥岭 [= Hainan, Baisha County,Yinggeling] / Ʋ坝水库 [= Hongba Reservoir] IJa [= lighting trap] / 2009.V.5 张ł [= Rui Zhang leg.] / 中LJnḿ物ñ [= IZAS]// IOZ(E)1948007–IOZ(E)1948006, IOZ(E)1948011’; 2♀♀ ( IZAS), ‘ 海南白沙县ƶ哥岭 [= Hainan, Baisha County, Yinggeling] / Ʋ坝水库 [= Hongba Reservoir] IJa [= lighting trap] / 2009.V.4 党ŨƲ [= Li-Hong Dang leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1948005, IOZ(E)1948010’; 2 ♀♀ ( IZAS), ‘ 海南白沙县ƶ哥 岭 [= Hainan, Baisha County, Yinggeling] / Ʋ坝水库 [= Hongba Reservoir] IJa [= lighting trap] / 2009.V.5 党ŨƲ [= Li-Hong Dang leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1948003–IOZ(E)1948004’; 2 JJ 3 ♀♀ ( IZAS), ‘ 海南$罗山Ṇ [= Hainan, top of Mount Diaoluoshan] / 海Ɲ： 900m / 2007.V.3 Ηκ [= Hui Xiao leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1947926, IOZ(E)1947930, IOZ(E)1947966, IOZ(E)1947979, IOZ(E)1947988’; 5 JJ 1 ♀ ( IZAS), ‘ 海南$罗山 [= Hainan, top of Mount Diaoluoshan] / 海Ɲ： 900m / 2007.V.4 Ηκ [= Hui Xiao leg.] / 中LJn ḿ 物ñ [= IZAS] // IOZ(E)1947931, IOZ(E)1947940, IOZ(E)1947965, IOZ(E)1947967, IOZ(E)1947981, IOZ(E)1947989’; 2JJ 13♀♀ ( IZAS), ‘ 海南$罗山Dz理ü [= Hainan, Management Station of Mount Diaoluoshan] / 2007.V.5 Ηκ [= Hui Xiao leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1947918, IOZ(E)1947927– IOZ(E)1947929, IOZ(E)1947932, IOZ(E)1947955, IOZ(E)1947959, IOZ(E)1947961, IOZ(E)1947963, IOZ(E)1947968, IOZ(E)1947983, IOZ(E)1947987, IOZ(E)1947990, IOZ(E)1947993–IOZ(E)1947994’; 3 JJ 1 ♀ ( IZAS), ‘ 海南$罗山⁂假村 [= Hainan, Mount Diaoluoshan, Vocational Village] / 海Ɲ： 920m / 2007.V.4 / 中LJnḿ物ñ [= IZAS] // 18.39°N / 108.53°E / 张彦周 [= Yan-Zhou Zhang leg.] / 中LJnḿ物 ñ [= IZAS] // IOZ(E)1947936–IOZ(E)1947937, IOZ(E)1947969, IOZ(E)1947980’; 2 JJ 14 ♀♀ ( IZAS), ‘ 海南$罗山 [= Hainan, Mount Diaoluoshan] / 海Ɲ： 260m / 2007.V.4-5 / 中LJnḿ物ñ [= IZAS] // 18.39°N / 108.53°E / 张彦周 [= Yan-Zhou Zhang leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1947933–IOZ(E)1947934, IOZ(E)1947938, IOZ(E)1947954, IOZ(E)1947956–IOZ(E)1947958, IOZ(E)1947960, IOZ(E)1947971, IOZ(E)1947974–IOZ(E)1947977, IOZ(E)1947991, IOZ(E)1947995, IOZ(E)1947999’; 8 JJ 13 ♀♀ ( IZAS), ‘ 海南五Dz山 [= Hainan, Mount Wuzhishan] / 海Ɲ： 730-900m / 2007.V.8-13 / 中LJ nḿ物ıffñ [= IZAS] // 18.54° N / 109.40° E / ze人：Ηκ 韩Ʋ Ɩ ṅt云 [= Xiao Hui, Hong-Xiang Han & Song-Yun Lang leg.] / 中 LJ n ḿ 物 ñ [= IZAS] // IOZ(E)1948606, IOZ(E)1947939, IOZ(E)1947941–IOZ(E)1947947, IOZ(E)1947950–IOZ(E)1947953, IOZ(E)1947964, IOZ(E)1947970, IOZ(E)1947972–IOZ(E)1947973, IOZ(E)1947982, IOZ(E)1947984–IOZ(E)1947986’; 1 ♀ ( IZAS), ‘ 海南 五Dz山水满乡 [= Hainan, Mount Wuzhishan, Shuimanxiang Township] / 度假寨 [= vocational village] 730m IJa [= lighting trap] / 2007.V.9 / 中LJnḿ物ıffñ [= IZAS] // 18.54° N / 109.40° E / 李娴 [= Xian Li leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1947948’; 2 ♀♀ ( IZAS), ‘ 海南五Dz山水满乡 [= Hainan, Mount Wuzhishan, Shuimanxiang Township] / 2007.V.8 李娴 [= Xian Li leg.] / 中LJnḿ物ıffñ [= IZAS] // IOZ(E)1947921, IOZ(E)1947949’; 5 JJ 18 ♀♀ ( IZAS), ‘ 海南五Dz 山 [= Hainan, Mount Wuzhishan] / 920m 2007.V.8 / 李娴 [= Xian Li leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1948012–IOZ(E)1948031, IOZ(E)1948033, IOZ(E)1948605, IOZ(E)1948688’; 5 JJ 1 ♀ ( IZAS), ‘ 海南五Dz山‼口 [= Hainan, gate of Mount Wuzhishan] / 海Ɲ： 708m / 2010.IV.8 / 中LJnḿ物ñ [= IZAS] // 18.90° N / 109.67° E / ze 人：hxẉ [= Mei-Ying Lin leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1948094, IOZ(E)1948095, IOZ(E)1948097, IOZ(E)1948098, IOZ(E)1948099, IOZ(E)1948101’; 8 JJ 1 ♀ ( IZAS), ‘ 海南五Dz山保 护区 [= Hainan, Mount Wuzhishan Nature Reserve] / 海Ɲ： 708m / 2010.IV.9 / 中LJnḿ物ñ [= IZAS] // 18.90° N / 109.67° E / ze人： hx ẉ [= Mei-Ying Lin leg.] / 中LJn ḿ物ñ [= IZAS] // IOZ(E)1948084–IOZ(E)1948085, IOZ(E)1948088–IOZ(E)1948093, IOZ(E)1948100’; 12JJ 25♀♀ ( IZAS), ‘ 海南五Dz山保护ü [= Hainan, Mount Wuzhishan] / 海Ɲ： 708m / 2010.IV.11 / 中LJnḿ物ıffñ [= IZAS] // 18.90° N / 109.67° E / ze人：hxẉ [= Mei-Ying Lin leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1948032, IOZ(E)1948036– IOZ(E)1948043, IOZ(E)1948119–IOZ(E)1948146’; 4 JJ ( IZAS), ‘ 海 南DZ江县NJƗ岭 [= Hainan, Changjiang County, Mount Bawangling] / 东二保护ü [= Dong’er Station] 825m / 2007.V.9 / 中LJnḿ物ñ [= IZAS] // N 19°12′/ E 109°10′ / 梁±l Ηκ [= Hong-Bin Liang & Hui Xiao leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1946843–IOZ(E)1946844, IOZ(E)1946847–IOZ(E)1946848’; 5JJ 6♀♀ ( IZAS), ‘ 海南尖峰岭ƾ 池 [= Hainan, Mount Jianfengling, Tianchi] / 海Ɲ： 675m / 2007.V.1-5 / 中LJnḿ物ñ [= IZAS] // N 18°71′/ E 108°87′ / 梁±l Ηκ [= Hong-Bin Liang & Hui Xiao leg.] / 中LJnḿ物ñ [= IZAS] // IOZ(E)1946845–IOZ(E)1946846, IOZ(E)1946849–IOZ(E)1946857’; 1 J ( ZMGC), ‘ 海南省五Dz山亚泰ffih酒店 [= Hainan Prov., Wuzhishan, Yatai Rainforest Hotel] / N 18.907457° E 109.676927°/ 720m, 2019.IV.8 周‖ [= Run Zhou leg.] / 中LJnḿ物ñ [= IZAS] // Ex. coll. IZCAS’; 1J 1♀ ( ZMGC), ‘ 海南白沙ƶ哥岭公×Ι [= Hainan, Baisha, Yinggeling, roadside] / 109°34′6.88″E 19°2′43.59″N / Ɨ兴民 佟※博 梁建 锋 [= Xing-Min Wang, Jun-Bo Tong & Jian-Feng Liang leg.] / 2019. IV.14 589m IJa法 [= light trap]’; 1 J ( ZMGC), ‘ 海南白沙ƶ哥岭 公×Ι [= Hainan, Baisha, Yinggeling, roadside] / 109°34′6.88″E 19°2′43.59″N / Ɨ兴民 佟※博 梁建锋 [= Xing-Min Wang, Jun-Bo Tong & Jian-Feng Liang leg.] / 2019.IV.13 589m IJa法 [= light trap]’; 1 J 3 ♀♀ (1 J 2 ♀♀ in NMPC, 1 ♀ in CZGC), ‘ CHINA, HAINAN Isl., 4-6.v.2011 / Limushan Mts. / frst administer. centre (at light) / 19°10′30″N, 109°44′33″E, 630m / M. Fikáček, V. Kubeček & L. Li leg.’; 3JJ ( NMPC), ‘ CHINA, HAINAN Isl., 7-8.v.2011 / Bawangling Mts. / BAOTIE env. (at light) / 19°05.3′N, 109°07.4′E; 415-800 m / M. Fikáček, V. Kubeček & L. Li leg.’; 6 JJ ( CZGC), ‘ China, HAYNAN′ Isl. / Wuzhi Shan Mts., / 18′53N-109′43E, / 20.02- 10.04.2001 / 1500m. Loc. Coll. lg.’; 1 ♀ ( JZSC), ‘ China: Hainan, Wuzhishan / alt. 930m, 18°54′09″N / 109°41′18″E, 8–11.VI.2017, / 8–11.VI.2017, / Jiang, Ge leg.’; 3 JJ ( WFCC), ‘ 2019.IV.12 ~14 / 海南五Dz山 [= Hainan, Mount Wuzhishan] / 1200m / Wȋ玥z [= Jian-Yue Qiu leg.]’; 1 J ( ZMGC), ‘ 广东Ḛ州aeŧ山 [= Guangdong Province, Huizhou City, Mount Xiangtoushan] / 2015.III.26-27, / 叶b海z [=Xin-Hai Ye leg.]’; 1 J ( GEEP), ‘Guangdong’; 81 JJ 120 ♀♀ (62 JJ 79 ♀♀ in ZMGC, 5 JJ 15 ♀♀ in SCAU, 10 JJ 20 ♀♀ in IZAS, 3 JJ 5 ♀♀ in WFPC, 1 J 1♀ in MSPC), ‘ CHINA: Guangdong Prov., / Maoming City, Xintian Town, / N 21°42′26.16″ E 111°17′3.28″/ light trap, Qiang GAN leg.’; 5 JJ ( CZGC), ‘ CHINA: Hainan Prov., Ledong, / Mts. Jianfengling, alt. 716m, / 18.710708 N 108.875812 E / 2019.IV, Shi-Liang Mo leg.’.

Description. Male (holotype). Body shape elongate ovoid, convex. Total length 18.0 mm, maximum width 9.1 mm.

Colour. Dorsal and ventral surfaces blackish-brown, with green metallic luster and coppery tinge; antennae, claws, teeth of protibia, margins of pronotum, and apex of scutellum reddish-brown.

Head. Clypeus semicircular, approximately 2.81 times wider than long, anterior margin reflexed, disc densely rugopunctate; frons densely punctate, each side with several larger punctures along eye, each of these large punctures with long seta; vertex finely punctate; ratio of interocular width/ width of head approximately 0.67; antennal club slightly longer than antennomeres II–VI combined.

Pronotum approximately 1.57 times wider than long; with smooth median longitudinal line, each side with depression; anterior angles acute and produced, posterior angles obtuse; sides almost straight and weakly converging anteriad in posterior three fifths, then fairly converging anteriad in anterior two fifths; basal marginal line interrupted before scutellum, all other marginal lines complete; surface densely and finely punctate; with several fine erect setae along lateral margins.

Scutellar shield. Sides rounded, apex acute; disc with sparse minute punctures.

Elytra ( Fig 3A ) regularly striate, strial punctures distinct; inner primary striae slightly impressed; subsutural interstice with two secondary striae, which unite towards apex, punctures irregularly doubled; whole surface moderately densely and finely punctate, punctures somewhat coalescent posterior to humerus; membranous apical border extending anteriad to level of front of metacoxa; epipleura with longitudinal row of short erect setae extending posteriad from level of metacoxa to level of abdominal ventrite III.

Legs. Protibia ( Fig. 4A ) approximately 3.5 times longer than wide; bidentate, proximal tooth blunt, apical tooth acute, extending to protarsomere II; protarsomere V ventrally with strong internomedial protruberance in middle; inner protarsal and outer mesotarsal claws incised apically, forming two branches; upper branches short, sharp at apex; lower branch of inner protarsal claw enlarged, sharp at apex; lower branch of outer mesotarsal claw slender, sharp at apex. Metatibia fusiform. Femora with transverse rows of moderately long erect setae; setae denser and longer on mesofemur.

Abdomen. Propygidium transversely striolate, striolation interrupted near pygidium. Pygidium triangular and moderately convex; rugostriolate, striolation denser on disc; sparsely surrounded by moderately long or long erect setae, setae denser on apex.

Ventral surface. Ventral prothoracic surface rugopunctate, bearing sparse long erect setae; ventral mesothoracic surface rugoso-punctate anteriorly, with irregular large punctures posteriorly, bearing dense short pubescence; metathoracic surface rugopunctate, bearing dense long pubescence, shallowly and sparsely punctate in middle. Abdominal ventrites I–IV carinate laterally; finely punctate, sides densely rugopunctate; with transverse band of sparse, long and semi-erect setae on posterior half (broadly interrupted in middle), dense and doubled laterad.

Male genitalia ( Figs 5A–C ). Parameres symmetrical, evenly curved inward sub-apically and pilose laterally, apices truncated and weakly emarginate ( Fig. 3 ).

Female. Protibia ( Fig. 4G ) reddish-brown; legs slenderer and shorter than in male, especially protarsus; apical tooth of protibia long, tongue-shaped, extending to protarsomere III; protarsomere V with indistinct internomedial protuberance; lower branch of inner protarsal claw thinner than in male; antennal club approximately as long as antennomeres II–VI combined.

Variation. Total body length and greatest width 14.5–18.7 mm and 7.3–9.1 mm (males), 17.4–18.4 mm and 8.7–9.9 mm (females), respectively. Body colour sometimes strongly coppery or reddish-brown. Ratio of pronotal width/length ranges from ca. 1.48 to 1.56. Punctures vary in size, those of inner primary and secondary striae sometimes more annulate, in extreme cases ca. 1.5 times wider than in the holotype. Some male specimens with shortened or abrased apical tooth of protibia ( Fig. 4C ), only extending to front of protarsomere I.

Differential diagnosis. Anomala consanguinea sp. nov. is closely allied to A. esmeralda Prokofiev, 2013 (central to southern Vietnam), A. russiventris Fairmaire, 1893 (major part of mainland Southeast Asia and southern China), and A. inepta Ohaus, 1916 (Malay Archipelago). These four species share a very similar general habitus and very similar male parameres with distinctly pilose sides. Anomala russiventris and A. inepta may be conspecific and represent a very widespread species (C. Zorn, pers. comm. 2018). The new species differs from A. esmeralda in having distinctly thicker protarsal claws and protarsomeres ( Figs 4A–C ). The apical tooth of protibia usually extends to front of protarsomere 2 in male and protarsomere 3 in female (slightly extending beyond the front of protarsomere 1 in male and protarsomere 2 in female in A. esmeralda ). Moreover, the parameres are strongly curved inward subapically and truncated apically ( Figs 5A–C ) (weakly curved inward subapically and acute apically in A. esmeralda , Figs 5D–F ). Anomala consanguinea sp. nov. can be distinguished from A. russiventris and A. inepta by sparser irregular punctures of elytra (moderately dense in A. consanguinea , Figs 3A–C , dense to very dense in A. russiventris and A. inepta , Figs 3E–F ), impressed inner primary striae of elytra (not impressed in A. russiventris and A. inepta ) and less wide pronotum (ca. 1.48 to 1.56 times wider than long in A. consanguinea , ca. 1.65 to 1.73 times wider than long in A. russiventris and A. inepta ). While most specimens of A. russiventris and A. inepta can be easily distinguished from A. consanguinea by brown ventral side, some A. russiventris specimens show the same dark metallic green color as found in the new species.

Etymology. The specific epithet is derived from the Latin adjective consanguineus, -a, -um (= related by blood), referring to the close relation between the new species and A. esmeralda .

Distribution. China (Hainan, Guangdong).

Figures

Fig. 1. Habitus of Anomala consanguinea sp. nov.A–C – holotype; D – paratype J (Guangdong, China); E – paratype J (Hainan, China); F – paratype ♀ (Hainan, China). A, D–F – dorsal view; B – lateral view; C – ventral view. Scale = 5 mm.

Fig. 3. Elytra of Anomala spp., male. A – A. consanguinea sp. nov., holotype; B – ditto, paratype (Guangdong, China); C – ditto, paratype (Hainan, China); D – A. esmeralda Prokofiev, 2013 (Quảng Nam, Vietnam); E – A. russiventris Fairmaire, 1893 (Guangdong, China); F – A. inepta Ohaus, 1916 (Sabah, Malaysia). Scale = 1 mm.

Fig. 4. Protarsi and protibial teeth of Anomala spp.A – A. consanguinea sp. nov.; holotype; B – ditto, paratypeJ (Guangdong, China); C – ditto, paratype J (Hainan, China); D – A. esmeralda Prokofiev, 2013 J (Quảng Nam,Vietnam); E – Anomala russiventris Fairmaire, 1893 J (Guangdong, China); F – A. inepta Ohaus, 1916 J (Sabah, Malaysia); G – A. consanguinea, paratype ♀ (Hainan, China). Scale = 1 mm.

Fig. 5. Aedeagi of Anomala spp.A–C – A. consanguinea sp. nov., holotype; D–F – A.esmeralda Prokofiev, 2013 (Quảng Nam,Vietnam); G–K – Anomala russiventris Fairmaire, 1893 (Guangdong, China); I–L – A. inepta Ohaus, 1916 (Sabah, Malaysia). A, D, G, J – frontal view; B, E, H, K – ventral view; C, F, I, L – lateral view. Sc 1: Scale = 1 mm for A–F; Sc 2: Scale = 1 mm for G–L.