Heptathela unten, Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang & Li, Daiqin, 2019

Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang & Li, Daiqin, 2019, A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands, ZooKeys 888, pp. 1-50 : 1

publication ID

https://dx.doi.org/10.3897/zookeys.888.34494

publication LSID

lsid:zoobank.org:pub:B995C056-97EC-41A4-9012-B58F9D3AFDC1

persistent identifier

https://treatment.plazi.org/id/65FF6B26-074B-4DC8-87F8-021268D23D87

taxon LSID

lsid:zoobank.org:act:65FF6B26-074B-4DC8-87F8-021268D23D87

treatment provided by

ZooKeys by Pensoft

scientific name

Heptathela unten
status

sp. nov.

Heptathela unten sp. nov. Figs 22 View Figure 22 , 23 View Figure 23

Type material.

Holotype: JAPAN · ♂; Okinawa-ken, Nakijin-son, Unten Port; 26.68N, 128.00E; alt. 25 m; 27 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-522.

Paratypes: JAPAN · 1 ♂, 2 ♀♀; same data as for holotype; XUX-2012-523, 527, 528A · 3 ♀♀; same data as for holotype; 10 May 2014; D. Li and B. Wu leg.; XUX-2014-083, 083A, 083B.

Diagnosis.

Males of H. unten sp. nov. can be distinguished from those of H. yanbaruensis by the blunt tegular marginal apophysis ( Fig. 22A, C, D View Figure 22 ); from those of H. helios by the conductor with serrated margin and small tegular marginal apophysis ( Fig. 22A, C View Figure 22 ). Females of H. unten sp. nov. cannot be distinguished morphologically form those of H. crypta sp. nov. ( Fig. 23 A–L View Figure 23 ). However, H. unten sp. nov. can be diagnosed from all other Okinawa group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: G (482), C (635).

Description.

Male (Holotype). Carapace yellow brown; opisthosoma brown, with dark brown tergites close to each other; cheliceral groove with ten denticles; seven spinnerets. Measurements: BL 10.00, CL 5.05, CW 4.60, OL 4.60, OW 2.90; ALE> PLE> PME> AME; leg I 15.71 (4.48 + 1.93 + 3.30 + 4.00 + 2.00), leg II 16.59 (4.45 + 1.90 + 3.37 + 4.62 + 2.25), leg III 18.29 (4.50 + 1.93 + 3.58 + 5.53 + 2.75), leg IV 23.40 (5.82 + 2.28 + 4.52 + 7.30 + 3.48).

Palp. Prolateral side of paracymbium unpigmented and unsclerotised, numerous setae and spines at the tip of paracymbium ( Fig. 22A, B View Figure 22 ). Contrategulum margin incurved nearly in the middle and the contrategulum divided into proximally serrated and distally smooth margins ( Fig. 22A, D View Figure 22 ). Tegulum with wide dorsal extension of terminal apophysis ( Fig. 22C, D View Figure 22 ), blunt terminal and small marginal apophysis ( Fig. 22A, C, D View Figure 22 ). Conductor sclerotised and ovate, prolateral conductor with one or two shallow folds, and with a serrated margin ( Fig. 22B, C View Figure 22 ). Embolus sclerotised, with a wide opening, the distal margin slightly sclerotised, and with a saddle-shaped margin in the retrolateral view ( Fig. 22B, C View Figure 22 ).

Females (N = 5). Carapace and opisthosoma colour as in male, dark brown tergites separated from each other; cheliceral groove with 12 or 13 pronounced denticles; seven or eight spinnerets. Measurements: BL 7.81-12.00, CL 3.60-4.55, CW 3.30-4.40, OL 4.10-7.60, OW 3.00-6.20; ALE> PLE> PME> AME; palp 6.64 (2.19 + 1.22 + 1.45 + 1.78), leg I 7.48 (2.47 + 1.35 + 1.22 + 1.52 + 0.92), leg II 7.40 (2.21 + 1.30 + 1.21 + 1.65 + 1.03), leg III 7.79 (2.11 + 1.38 + 1.20 + 1.90 + 1.20), leg IV 11.55 (3.20 + 1.55 + 2.10 + 3.05 + 1.65).

Female genitalia. A pair of depressions on the ventro-lateral part of genital atrium indistinct ( Fig. 23D, E View Figure 23 ). Paired receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, the inners similar or smaller than the laterals, paired receptacular clusters tuberculate, inners with or without genital stalks ( Fig. 23A, B, D, E View Figure 23 ).

Etymology.

The species epithet, a noun in apposition, refers to the type locality, Unten Port.

Distribution.

The species is endemic to the Japanese island Okinawajima ( Fig. 1C View Figure 1 ).

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Liphistiidae

Genus

Heptathela