Cylindera (Cylinderina) werneri Wiesner, 1988

Wiesner, Jürgen & Dheurle, Charles, 2018, Remarks on the subgenus Cylinderina Rivalier (Coleoptera: Cicindelidae: Cylindera) from the Philippines with description of a new species 144. Contribution towards the knowledge of the Cicindelidae, Insecta Mundi 670, pp. 1-16 : 7

5. Cylindera (Cylinderina) werneri Wiesner

( Fig. 29 )

Cylindera (Cylinderina) werneri Wiesner 1988: 153 , Fig. 1–5 .

Cylindera (Cylinderina) werneri . Wiesner 1992: 187; Cabras et al. 2016: 196; Dheurle 2016: 39, Fig. 5, 9 .

Type depository. Holotype male in ZSMC, paratype female in JWGC.

Type status. Holotype male in ZSMC, type labels: “Queson National, Forest Park, Luzon 24. March. 1981, coll. N. Nishikawa; Typus [red, printed]; ZSM - Samml ., K. Werner 2009; HOLOTYPUS, CYLIN- DERA, ( Cylinderina) werneri Wiesner [type written, red]”.

Paratype female in JWGC, type labels: “Queson National, Forest Park, Luzon 24. March. 1981, coll. N. Nishikawa; PARATYPUS, CYLINDERA , ( Cylinderina) werneri Wiesner [type written, red]” .

Other material examined. 1 male in SDEI: “ Quezon Park, Tayabas, Luzon, P.I. VI-1-32; F. C. Hadden, collector” . 1 female in SDEI with same labels, but VI-6-32. 1 male in SDEI with same labels, but VI-9-32 .

Distribution. Philippines, Luzon (Prov. Quezon, National Forest Park, Quezon Park).

Diagnosis. Distinguished by the reddish femora, the curved metallic elytral margin, and lacking subapical dot.

Re-description. Size: Total length (without labrum) 5.7–6.4 mm (mean = 5.9 mm, n = 5). Head: Dark bluish green above, genae dark blue, surface smooth, glabrous, with each two setigerous punctures next to the eyes in front and at the center; strong longitudinal striae on vertex and orbital plates, the striae becoming slightly concentric on vertex, and finer, irregular, transversely waved, behind on neck; genae roughly longitudinally striated. Eyes pale yellow. Ratio between width of head and elytra = 1.1 in medium. Labrum (male Fig. 33 , female Fig. 34 ) distinctly wider than long, ratio between width and length = 2.2 in medium; with six marginal setae, unidentate in the middle of front edge; color rufous, at base and lateral margins with blue green hue in various expansion. Mandibles yellowish, brownish on apical and inner teeth in males and completely brownish except for the base in females. Labial and maxillary palpi testaceous, the last joint metallic dark; basal segment of labial palpi with long erected white setae. Antennae slender, long, reaching half of the elytral length in the males, a little shorter in the females; scape with one long apical seta; antennomeres one to four brownish with metallic luster; antennomeres five to eleven dark blackish brown, dull, finely and evenly pubescent. Thorax: Pronotum as wide as long (ratio between width and length = 1.0 in medium), sides somewhat rounded between the transversal constrictions; glabrous, with some white setae laterally before the anterior constriction; transversely waved before and behind the transversal constrictions and at the lateral margin, with irregular wrinkles on disc; color bluish green. Sternae and episternae brownish black with bluish hue, proepisternum with several white setae at the front margin, mesepisternum glabrous, the remaining setose; mesepisternum of females with a longitudinal furrow over its whole length. Elytra: ( Fig. 30–32 ) parallel-sided; color dull brown on the disk, broadly and curved bordered from the scutellum up to the apex by blue greenish color; elytral testaceous maculation lost, or consists of a tiny subhumeral dot, or of a small subhumeral dot and a thin middle band. Surface with a fine microsculpture above, margin- ally smooth, shoulders glossy; preapical impression strong, apical microserration extremely fine; apical border distinctly rounded and restricted towards middle suture, with a small sutural spine. Epipleurae rufous. Ventral aspect: Venter setose apically, brownish black; coxae with white setae, testaceous; trochanters glabrous and testaceous; femora testaceous, brownish towards apex; tibiae and tarsi dark brownish; legs covered with some white setae, mesotibiae more densely setose. Aedeagus: ( Fig. 35 ) on left lateral view fusiform, straight, tapering, slightly enlarged in the middle, with a straight, blunt and slightly upturned apex (total length 2.1mm).

Remarks. While describing C. (C.) werneri , the first author overlooked a tiny subhumeral dot on the elytra of the male holotype.

Figures

Figure 1. Body length of Cylindera (Cylinderina) species in mm (without labrum).

Figure 2. Distribution map of Cylindera (Cylinderina) species.

Figures 3–16. Cylindera (Cylinderina). All scales = 1 mm. 3–10. C. (C.) nanula (Horn, 1937). 3) Habitus, paralectotype male from Paete (SDEI). 4–6. Left elytron. 4) Paralectotype male from Paete (SDEI). 5) Female from Santa Ana (JWGC). 6) Female from Mt. Province (JWGC). 7–9. Labrum. 7) Paralectotype male from Paete (SDEI). 8) Female from Santa Ana (JWGC). 9) Female from Mt. Province (JWGC). 10) Left lateral view of aedeagus, paralectotype from Paete (SDEI). 11–16. C. (C.) nana (Schaum, 1862). 11) Habitus, male from Kayapa (CDFC). 12–13. Left elytron. 12) Male from Kayapa (CDFC). 13) Female from Mt. Province (JWGC). 14) Left lateral view of aedeagus, male from Kayapa (CDFC). 15–16. Labrum. 15) Male from Kayapa (CDFC). 16) Female from Mt. Province (JWGC).

Figures 29–41. Cylindera (Cylinderina). All scales = 1 mm. 29–35. C. (C.) werneri Wiesner, 1988. 29) Habitus, holotype male from National Forest Park (ZSMC). 30–32. Left elytron. 30) Holotype male from National Forest Park (ZSMC). 31) Male from Quezon Park (SDEI). 32) Paratype female from National Forest Park (JWGC). 33–34. Labrum. 33) Holotype male from National Forest Park (ZSMC). 34) Paratype female from National Forest Park (JWGC). 35) Left lateral view of aedeagus, male from Quezon Park (SDEI). 36–41. C. (C.) rothschildi (Horn, 1896). 36) Habitus, paratype male from Albay (SDEI). 37–38. Left elytron. 37) Paratype male from Albay (SDEI). 38) Paratype female (SDEI). 39) Left lateral view of aedeagus, paratype from Albay (SDEI). 40–41. Labrum. 40) Paratype male from Albay (SDEI). 41) Paratype female (SDEI).