Cylindera (Cylinderina) nanula (Horn)

Wiesner, Jürgen & Dheurle, Charles, 2018, Remarks on the subgenus Cylinderina Rivalier (Coleoptera: Cicindelidae: Cylindera) from the Philippines with description of a new species 144. Contribution towards the knowledge of the Cicindelidae, Insecta Mundi 670, pp. 1-16 : 2-3

1. Cylindera (Cylinderina) nanula (Horn)

( Fig. 3 )

Cicindela nanula Horn 1937: 57 .

Cylindera (Jansenia) nanula . Schilder 1953: 548.

Cylindera (Cylinderina) nanula . Wiesner 1992: 186; Cabras et al. 2016: 196.

Type depository. Lectotype male and paralectotype male in SDEI (here designated).

Type status. Lectotype male in SDEI, type labels: “LUZON, Laguna Pr, Mc Gregor, Paete ; Type, W.

Horn; Mus. Wash. [hand written]; SYNTYPUS [printed, red]; Dr. W. Horn det. 1935, C. nanula W. H., type [partially hand written]; nanula m. [pale yellow, hand written, black bordered]; Coll. W. Horn, Eberswalde; SDEI Coleoptera , # 301416; LECTOTYPE, Cylindera (Cylinderina) nanula (W. Horn, 1937) , ded. J. Wiesner, 2018”.

Paralectotype male in SDEI, type labels: “LUZON, Laguna Pr, Mc Gregor, Paete; Mus. Wash. [hand written]; Type, W. Horn; SYNTYPUS [printed, red]; Coll. W. Horn, Eberswalde; PARALECTOTYPE, Cylindera (Cylinderina) , nanula (W. Horn, 1937) , ded. J. Wiesner, 2018” .

Other material examined. 1 male in SDEI: “ Laguna, Ins. Phil .” . 1 female in NHMW: “ Philippinen, Luzon, Imugan / S. Drechsler , 6. 1917” . 1 female in JWGC: “ PHILIPPINES, Mt. Province, N. Luzon, VI. 1988 ” . 1 female in CDFC: “ PHILIPPINES, North Luzon, Cagayan, Sta Ana / VIII. 2015, LUMAWIG leg.” . 2 males, 1 female in CKGC: “Maddela town env. – Luzon, Sierra Madre – PHILIPPINES, local collector 04/2018” .

Distribution. Philippines, Luzon (Prov. Cagayan: Santa Ana, Prov. Laguna: Paete, Prov. Mt. Province, Prov. Nueva Vizcaya: Imugan, Prov. Quirino: Maddela).

Diagnosis. Distinguished from all other members of the subgenus by the elytral maculation.

Re-description. Size: Total length (without labrum) 5.8–7.0mm (mean = 6.2mm, n = 6). Head: Coppery above, metallic greenish on clypeus, frons, orbital plates, and genae; surface smooth, glabrous, with each two setigerous punctures next to the eyes in front and at the center; strong longitudinal striae on vertex and orbital plates, the striae becoming slightly concentric on vertex, and finer, irregular, transversely waved, behind on neck; genae roughly longitudinally striated. Eyes pale yellow. Ratio between width of head and elytra = 1.2 in medium. Labrum (male Fig. 7 , female Fig. 8, 9 ) distinctly wider than long, ratio between width and length = 2.0 in medium; with five to seven marginal setae, unidentate in the middle of front edge; color in various extension metallic greenish at lateral and basal margin, the remainder part of the labrum brownish yellow, at least a small area around the marginal tooth. Mandibles yellowish, more or less rufescent or darkened on apical and inner teeth. Labial and maxillary palpi testaceous, the last joint metallic dark greenish; basal segment of labial palpi with long erected white setae. Antennae slender, long, reaching the apical third of the elytral length in the males, the half of the elytral length in the females; scape brownish yellow, with one long apical seta; antennomeres two to four mostly glabrous, dark, with metallic luster, sometimes rufous at apical end; antennomeres five to eleven dark blackish brown, dull, finely and evenly pubescent. Thorax: Pronotum as wide as long (ratio between width and length = 1.0 in medium), straight at sides, glabrous, transversely waved before and behind the transversal constrictions and at the lateral margin, with irregular wrinkles on disc; color coppery above, bluish green at lateral margin. Sternae and episternae metallic dark greenish or dark coppery, almost glabrous, mesepimeron and metasternum with some white setae ventrally, metepisternum with some white setae at base; mesepisternum of females with a longitudinal furrow over its whole length. Elytra: ( Fig. 4–6 ) parallel-sided, somewhat divergent in males, somewhat rounded in females; color dull brown on the disk near the suture, apically lightened in some specimens, laterally in variable extension bluish green; elytral testaceous maculation consists of a slender subhumeral lunule, a broader middle band that nearly reaches the middle suture, and a large subapical dot. Surface with a fine microsculp- ture, and numerous larger, evenly spaced punctures laterally, shoulders glossy; preapical impression strong, apical microserration extremely fine; apical border distinctly rounded and restricted towards middle suture, with a small sutural spine. Epipleurae rufous. Ventral aspect: Venter dark brown, with some thin white setae marginally; coxae with white setae; coxae and trochanters testaceous; femora testaceous, apical tip (knee) darkened; tibiae and tarsi dark, with little metallic reflections; legs covered with some with setae, mesotibiae more densely setose. Aedeagus: ( Fig. 10 ) on left lateral view fusiform, stout and straight, tapering, slightly enlarged at the front and the back, with a short, straight, blunt apex (total length 2.2mm).

Remarks. Apparently C. (C.) nanula lives in several biotopes throughout Luzon. Nevertheless, more detailed collecting data would be welcome to make the occurrence of this species more precise.

Figures

Figures 3–16. Cylindera (Cylinderina). All scales = 1 mm. 3–10. C. (C.) nanula (Horn, 1937). 3) Habitus, paralectotype male from Paete (SDEI). 4–6. Left elytron. 4) Paralectotype male from Paete (SDEI). 5) Female from Santa Ana (JWGC). 6) Female from Mt. Province (JWGC). 7–9. Labrum. 7) Paralectotype male from Paete (SDEI). 8) Female from Santa Ana (JWGC). 9) Female from Mt. Province (JWGC). 10) Left lateral view of aedeagus, paralectotype from Paete (SDEI). 11–16. C. (C.) nana (Schaum, 1862). 11) Habitus, male from Kayapa (CDFC). 12–13. Left elytron. 12) Male from Kayapa (CDFC). 13) Female from Mt. Province (JWGC). 14) Left lateral view of aedeagus, male from Kayapa (CDFC). 15–16. Labrum. 15) Male from Kayapa (CDFC). 16) Female from Mt. Province (JWGC).