Mycomya goral , Väisänen, Rauno, 2013

Väisänen, Rauno, 2013, New Mycomya species from the Himalayas (Diptera, Mycetophilidae): 3. Subgenera Cesamya and Mycomyopsis, Zootaxa 3737 (2), pp. 129-153: 133-137

publication ID

http://dx.doi.org/10.11646/zootaxa.3737.2.3

publication LSID

lsid:zoobank.org:pub:358CB400-2CF2-4AF1-8DC6-9F8C57721C33

persistent identifier

http://treatment.plazi.org/id/600087D2-FFA5-4834-5086-FD7EFEF61CD4

treatment provided by

Plazi

scientific name

Mycomya goral
status

sp. n.

Mycomya goral  sp. n.

Figs. 2View FIGURE 2 A –F

Material studied. Holotype. ♂. NEPAL, Ktmd. Godavari, 6000 ', 17.VII. 1967, Can Nepal Exp. (in CNC). Paratypes (only ♂♂). Same data, 1 ♀ (in CNC); same data, but Mal. tr., 21.VII. 1967, 1 ♂, 1 ♀ (in CNC); same data, but Mal. tr., 2.VIII. 1967, 2 ♂♂ (in CNC); same data, but 3.VIII. 1967, 2 ♂♂ (in CNC and MZH).

Description. ♂. Head. Palp yellow, basally yellowish to brownish, other mouthparts yellow, face yellow, posterior parts of head brown. Antenna brownish, scape, pedicel and base of 1 st flagellomere yellow. Length of 1 st flagellomere about 3.5 x, 2 nd flagellomere 2 x, its width. Thorax. Pronotum yellow, with 2 long setae. Scutum yellowish to brownish with 3 indistinct brownish to brown longitudinal, posteriorly fused stripes, anterolateral corners of scutum yellow. Anepisternum yellow. Preepisternum yellow. Scutellum light brownish, with 4 long setae. Laterotergite light brownish. Mediotergite light brownish, bare. Wing. Length 3.2–3.5 mm. Wing hyaline. Sc ending in R 1 near or distad of middle of small cell, Sc 1 apically broken or entirely missing. Apical part of Sc bearing 6–13 small setae. Small cell 1– 2 x as long as wide. Cu fork distinctly distal to M fork. M ratios: 0.95–1.05, 1.38–1.54. Cu ratios: 0.84–0.95, 1.40–1.67. Small setae: M petiole: 0; M 1: 0–5; M 2: 0–1; Cu petiole: 0; Cu 1: 0; Cu 2: 0. Halter pale yellowish, apex yellowish to brownish. Legs. Coxae yellow, femora yellow, tibiae and tarsi brownish to brown. Coxa 2 without spur. Leg ratios: bt 1:t 1 = 0.74–0.75, bt 2:t 2 = 0.63–0.67, bt 3:t 3 = 0.56–0.60. Abdomen. Tergites brownish to brown, sternites slightly paler. Hypopygium. Figs. 2View FIGURE 2 A –F, yellowish. Tergal part with 1 wide median comb of short spines, and 2 narrow inner combs, both with about 5 spines. Dark inner spur of tergal part apically rounded ( Fig. 2View FIGURE 2 C). Tergal lateral appendage setose, slender, about 5 x as long as wide, with 2 flattened, curved subapical setae ( Fig. 2View FIGURE 2 D). Sternal submedian filament long, slender, curved, extending far beyond apex of aedeagus. Gonostylus about 5 x as long as wide, with 3 subapical teeth, 1 strong seta, and membranous lateral lobe more than ½ of length of main branch of gonostylus ( Fig. 2View FIGURE 2 E). Aedeagus with 2 lateral lobes, extending slightly beyond its apex ( Fig. 2View FIGURE 2 F). Female. Wing length 4.0 mm. Thorax yellow, scutum with brown medial stripe, yellowish submedial stripes and brownish sublateral stripes, anterolateral corners of scutum yellow, mediotergite and laterotergite light brownish. Abdomen light brownish. Leg ratios: bt 1:t 1 = 0.83, bt 2:t 2 = 0.67, bt 3:t 3 = 0.60. Terminalia yellow.

Discussion. Mycomya goral  is a small species and closely related to M. niltava  sp. n. and M. sachak  sp. n., both of which have a similar wide tergal comb of small spines, a similar general shape of the tergal lateral appendage ( Figs. 2View FIGURE 2 D, 3 E, 4 D), and long curved sternal submedian filaments. Mycomya goral  differs from M. sachak  in having an isolated flattened spine in the middle part on the inner margin of the tergal lateral appendage ( Fig. 2View FIGURE 2 D) similar to that in M. niltava  ( Fig. 3View FIGURE 3 E). See also discussion under M. niltava  .

Etymology. The species name is a noun in apposition and refers to the Himalayan goral  , Nemorhaedus goral (Hardwicke)  , a bovid species found across the Himalayas.

Mycomya niltava  sp. n. Figs. 3View FIGURE 3 A –G

Material studied. Holotype. ♂. NEPAL, Dobang Kharka, 2400 m 83 ° 24 'E. 28 ° 36 'N. Oct. 28 1971 A. Nakanishi (in KUC). Paratypes (only ♂♂). Same data as holotype, 2 ♀♀ (in KUC); same data, but 9.X. 1971, 4 ♂♂, 6 ♀♀ (in KUC); same data, but 19.X. 1971, 1 ♀ (in KUC); same data, but 20.X. 1971, 28 ♂♂, 79 ♀♀ (in KUC); same data, but 23.X. 1971, 1 ♂, 1 ♀ (in KUC); same data, but 30.X. 1971, 1 ♂ (in KUC); Lipshiba Kharka, 83 ° 23 'E, 28 ° 35 'N, 1900 m, 30.X. 1971, A. Nakanishi, 1 ♀ (in KUC).

Description. ♂. Head. Palp yellowish, other mouthparts and face brownish, posterior parts of head brown. Antenna brownish, scape, pedicel and base of 1 st flagellomere yellow. Length of 1 st flagellomere about 4 x, 2 nd flagellomere 2.5 x, its width. Thorax. Pronotum, with 3 long setae. Scutum yellowish with 3 brownish longitudinal stripes. Anepisternum yellowish to brownish. Preepisternum brownish, dorsal part lighter. Scutellum brownish, with 4 long setae. Laterotergite brownish, dorsal part lighter. Mediotergite brownish, bare. Wing. Length 3.9–4.3 mm. Wing hyaline. Sc ending in R 1 near or distad of middle of small cell, Sc 1 missing or very short. Apical part of Sc bearing 4–15 small setae. Small cell 1–1.5 x as long as wide. Cu fork distinctly distal to M fork. M ratios: 0.85– 1.04, 1.15–1.41. Cu ratios: 0.89 –1.00, 1.47–1.80. Small setae: M petiole: 0; M 1: 0; M 2: 0–1; Cu petiole: 0; Cu 1: 0; Cu 2: 0. Halter pale yellowish, apex brownish. Legs. Coxae and femora yellow, tibiae and tarsi brownish to brown. Coxa 2 without spur. Leg ratios: bt 1:t 1 = 0.77–0.83, bt 2:t 2 = 0.67–0.71, bt 3:t 3 = 0.57–0.64. Abdomen. Tergites brownish to brown, sternites yellowish. Hypopygium. Figs. 3View FIGURE 3 A –G, yellowish. Tergal part with 1 wide outer comb of spines, and 2 inner combs, both with about 7 spines. Almost whole base of outer comb widely setose, only its narrow median part bare. Dark inner spur of tergal part apically pointed ( Fig. 3View FIGURE 3 D). Tergal lateral appendage setose, slender, curved, about 6 x as long as wide, apically slightly wider, with 1 flattened seta on median side of middle part and some flattened, curved apical setae ( Fig. 3View FIGURE 3 E). Gonostylus very long and slender, at least 8 x as long as wide, with 2–3 subapical teeth and small membranous lateral lobe of 1 / 3 of length of main branch of gonostylus ( Fig. 3View FIGURE 3 F). Sternal submedian filament long and curved, extending far beyond apex of aedeagus. Aedeagus with 2 lobes, both with curved, spur-like group of setae. Female. Wing length 4.0–5.0 mm. Thorax as in ♂. Abdomen brownish to brown, sternites lighter, posteriorly yellow. Leg ratios: bt 1:t 1 = 0.77–0.81, bt 2:t 2 = 0.69–0.73, bt 3:t 3 = 0.53–0.63. Terminalia yellow.

Discussion. Mycomya niltava  has a conspicuous, isolated and flattened seta on the inner margin of its tergal lateral appendage ( Fig. 3View FIGURE 3 E) resembling that of M. goral  sp. n., which is a smaller species having a short gonostylus with a large lateral lobe ( Fig. 2View FIGURE 2 E), and a rounded apex of the tergal spur ( Fig. 2View FIGURE 2 C). Mycomya niltava  also differs from M. sachak  on the basis of the same two characteristics. The base of the wide tergal comb of M. niltava  is widely setose ( Fig. 3View FIGURE 3 A), whereas its medial part is widely bare in M. sachak  sp. n. and M. goral  sp. n. The hypopygium of M. niltava  generally resembles also that of M. sinica Yang  & Wu, 1989, but differs from it e.g. in the shape of the tergal lateral appendage and that of the gonostylus. The tergal lateral appendage of M. sinica  is broad and apically pointed and its gonostylus has a long membranous lobe.

Etymology. The species name is a noun in apposition and refers to the bird genus Niltava  Hodgson in the family of Old World flycatchers, Muscicapidae  .

Mycomya sachak  sp. n. Figs. 4View FIGURE 4 A –F

Material studied. Holotype. ♂. NEPAL, Bogara, 1700–2200 m, 83 ° 23 'E. 28 ° 35 'N. Malaise trap Sept. 20–21. 1971 A. Nakanishi (in KUC). Same data as holotype, but 19–20. IX. 1971, 2 ♂♂ (in KUC); same data, but 21–22. IX. 1971, 1 ♂ (in KUC); same data but not by Malaise trap, 21.IX. 1971, 3 ♂♂, 9 ♀♀ (in KUC).

Description. ♂. Head. Palp, other mouthparts and face yellow, posterior parts of head brown. Antenna brownish, scape, pedicel and base of 1 st flagellomere yellow. Length of 1 st flagellomere about 3.5 x, 2 nd flagellomere 2 x, its width. Thorax. Pronotum yellow, with 3–4 long setae. Scutum yellowish with 3 brown, fused, longitudinal stripes. Anepisternum yellowish to brownish. Preepisternum yellowish, ventral half brownish. Scutellum brownish, with 4 long setae. Laterotergite brownish. Mediotergite brownish, bare. Wing. Length 3.3–3.7 mm. Wing hyaline. Sc ending in R 1 near middle of small cell, Sc 1 missing. Apical part of Sc bearing 7–13 small setae. Small cell 1– 2 x as long as wide. Cu fork distal to M fork. M ratios: 0.94–1.17, 1.26.– 1.63. Cu ratios: 0.84– 0.85, 1.37–1.45. Small setae: M petiole: 0; M 1: 0; M 2: 0; Cu petiole: 0; Cu 1: 0; Cu 2: 0. Halter pale yellowish, apex brownish. Legs. Coxae and femora yellow, tibiae and tarsi brownish to brown. Coxa 2 without spur. Leg ratios: bt 1:t 1 = 0.71–0.77, bt 2:t 2 = 0.64–0.66, bt 3:t 3 = 0.57–0.59. Abdomen. Brownish. Hypopygium. Figs. 4View FIGURE 4 A –F: yellowish. Tergal part with 1 wide outer comb of spines and 2 very narrow inner combs, both with about 2 spines.

Only lateral parts of base of outer comb with some setae, median part widely bare. Dark inner spur of tergal part apically slightly rounded ( Fig. 4View FIGURE 4 C). Tergal lateral appendage setose, relatively short, slender, less than 4 x as long as wide, only slightly curved, with some flattened, curved apical-subapical setae ( Fig. 4View FIGURE 4 D). Gonostylus straight, more than 4 x as long as wide, with 2 teeth and large membranous lateral lobe slightly shorter than main branch of gonostylus ( Fig. 4View FIGURE 4 E). Sternal submedian filament very long, curved, extending far beyond apex of aedeagus. Aedeagus with some subapical curved setae on inner side ( Fig. 4View FIGURE 4 F). Female. Wing length 3.7 –4.0 mm. Thorax as in ♂. Abdomen brownish, sternites 1–5 yellowish. Leg ratios: bt 1:t 1 = 0.77–0.78, bt 2:t 2 = 0.66–0.70, bt 3:t 3 = 0.61–0.62. Terminalia yellow.

Discussion. Mycomya sachak  resembles M. goral  sp. n., but differs from it in its slightly larger size and in the absence of the isolated strong flattened seta in the middle of the inner margin of the tergal lateral appendage. The lateral appendage has several apical and subapical flattened curved setae close to each other ( Fig. 4View FIGURE 4 D). The tergal lateral appendage of M. sachak  is wider than M. goral  ( Fig. 2View FIGURE 2 D), and its subapical setae are shorter than the basal setae, whereas there is no clear difference in their respective lengths in M. goral  .

Etymology. The species name is a noun in apposition and refers to the name of the snow leopard in the Tibetan Bhotia language.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Diptera

Family

Mycetophilidae

Genus

Mycomya