Mycomyopsis Väisänen, 1984

Väisänen, Rauno, 2013, New Mycomya species from the Himalayas (Diptera, Mycetophilidae): 3. Subgenera Cesamya and Mycomyopsis, Zootaxa 3737 (2), pp. 129-153 : 137-138

Subgenus Mycomyopsis Väisänen, 1984

The male hypopygium of subgenus Mycomyopsis is characterized by the presence of a parallel pair of medial combs of short setae in the tergal part of the hypopygium, and another, less visible inner pair of combs; distinct setose tergal lateral appendage on both sides of the outer comb; a deeply bilobed sternal synsclerite with or without sternal submedian filaments, i.e. a long, narrow extensions stemming from the basal, medial parts of both the lobes of the sternal synclerite; an absence of sternal lateral appendages; and the gonostylus with a few strong apicalsubapical teeth and a membranous lateral lobe. Mycomyopsis lacks the male mid-coxal spur, has a bare mediotergite, and the vein Sc ends in R1. The abdominal tergite 8 is narrow, arcuate, medially slightly narrower, with several setae along the whole posterior margin (e.g. Figs. 5 G, 9F, 12G, 13G), the medial part is seldom bare ( Fig. 15 E). The hypogynal valve of the female terminalia is tapering towards its apex and has at least some strong, long setae, and the apical segment of cercus is oval (see Väisänen 1984: e.g. figs. 975, 992, 1012, 1028).

Mycomyopsis is a species-rich and almost cosmopolitan subgenus (Väisänen 1984), which also appears to be well-represented in the Himalayas.

Mycomya aix sp. n. Figs. 5 A–E

Material studied. Holotype. ♂. MYANMAR, N.E. Burma, Kambaiti, 2000 m, 4.VI. 1934, Malaise (in MZH).

Description. ♂. Head. Palp brownish, other mouthparts and face yellowish, posterior parts of head brown. Antenna brownish, scape, pedicel and base of 1st flagellomere yellow. Length of 1st flagellomere about 4x, 2nd flagellomere 2.5x, its width. Thorax. Pronotum yellow, with 3 long setae. Scutum anteriorly yellow, posteriorly darker, with 3 brownish to brown longitudinal, posteriorly fused stripes. Anepisternum yellow. Preepisternum yellow, ventral half brownish. Scutellum brownish, with 4 long setae. Laterotergite brownish. Mediotergite brownish, bare. Wing. Length 4.4 mm. Wing hyaline. Sc ending in R1 near middle of small cell, Sc1 missing. Apical part of Sc bearing no small setae. Small cell about 1.5x as long as wide. Cu fork distal to M fork. M ratios: 0.93, 1.19. Cu ratios: 0.85, 1.41. Small setae: M petiole: 0; M1: 0; M2: 0; Cu petiole: 0; Cu1: 0; Cu2: 0. Halter apically brownish. Legs. Coxae and femora yellow, tibiae and tarsi brownish to brown. Coxa 2 without spur. Leg ratios: bt1:t1 =?, bt2:t2 = 0.75, bt3:t3 =?. Abdomen. Tergites 1–5 light brownish, 6–7 brown, sternites slightly paler. Hypopygium. Figs. 5 A–F: yellowish. Tergal part with two pairs of combs. Outer tergal combs very wide, both consisting of about 15 strong cone-like, apically blunt spines; several similar cones also behind outer combs; inner combs both with about 7 spines ( Fig. 5 C). Tergal lateral appendage very broad, slightly longer than its width, basally narrower, densely setose, with dark spur that is formed by strong, curved setae on medial side ( Fig. 5 D; see also Fig. 5 A, on both sides of outer combs). Sternal submedian filament missing ( Fig. 5 B, lower medial parts of deeply bilobed sternal synsclerite without filament-like extensions). Gonostylus about 5x as long as wide, with 6 subapical teeth and long membranous lobe, extending slightly beyond apex of gonostylus ( Fig. 5 E). Aedeagus comparatively long, slender, with relatively long lateral appendages on both sides almost reaching its apex ( Fig. 5 F, mid-upper inner part, between gonostyli).

Discussion. The complex structure of the tergal combs consisting of strong dark cone-like spines ( Figs. 5 A, 5C) and the dense setosity with the dark spur-like group of curved setae on the inner margin of the tergal lateral appendage ( Fig. 5 D) of M. aix are not known among the other species of the subgenus Mycomyopsis . Also the short wide and densely setose tergal lateral appendage is diagnostic.

Etymology. The species name is a noun in apposition and refers to the genus of the mandarin duck, Aix F. Boie.

Figures

FIGURE 5. Mycomya aix sp. n., male hypopygium. A – B tergal and sternal views, C. detail of tergal combs, twice enlarged, D tergal lateral appendage, twice enlarged, E gonostylus, twice enlarged, F apex of aedeagus, lateral view, twice enlarged G tergite 8. Scale 0.2 mm.

FIGURE 15. Mycomya unipectinata Edwards, male hypopygium. A – B tergal and sternal views, C tergal lateral appendage, twice enlarged, D gonostylus, twice enlarged, E tergite 8. Scale 0.2 mm.