Coccygidium Saussure, 1892

Sharkey, Michael, Yu, Dicky, van Noort, Simon, Seltmann, Katja & Penev, Lyubomir, 2009, Revision of the Oriental genera of Agathidinae (Hymenoptera, Braconidae) with an emphasis on Thailand and interactive keys to genera published in three different formats, ZooKeys 21 (2), pp. 19-54 : 36-37

Coccygidium Saussure, 1892

Type species: Coccygidium luteum Saussure, 1892 .

Brachyropalum Kriechbaumer, 1894 , first synonymized by Chou and Sharkey (1989) and confirmed by Sharkey (1996) and Sarmiento and Sharkey (2005). Type species: Brachyropalum pallidum Kriechbaumer, 1894 .

Brachyrhopalum Dalla Torre, 1898, synonymized by Chou and Sharkey (1989). Emendation for Brachyropalum .

Neophylax Ashmead, 1900 , first synonymized by Chou and Sharkey (1989) and confirmed by Sharkey (1996) and Sarmiento and Sharkey (2005). Type species: Neophylax snyderi Ashmead, 1900 .

Ahngeria Kokujev, 1902 , first synonymized by van Achterberg and Maeto (1990) and confirmed by Sharkey (1996, 1998) and Sarmiento and Sharkey (2005). Type species: Ahngeria transcaspica Kokujev, 1902 .

Lisitheria Cameron, 1904 , first synonymized by Chou and Sharkey (1989) and confirmed by Sharkey (1996); Sarmiento and Sharkey (2005). Type species: Lisitheria nigricornis Cameron, 1904 .

Xanthomicrodus Cameron, 1904 , first synonymized by Chou and Sharkey (1989) and confirmed by Sharkey (1996) and Sarmiento and Sharkey (2005). Type species: Xanthomicrodus iridipennis Cameron, 1904 .

Caenophylax Schulz, 1911 , first synonymized by Chou and Sharkey (1989) and confirmed by Sharkey (1996) and Sarmiento and Sharkey (2005). New name for primary homonym Neophylax Ashmead, 1900 , nec Neophylax McLachlan, 1871 .

Distribution: Oriental, Palaearctic, Oceanic, Australian, African, primarily tropical and warm-temperate, but reasonably represented in moderate temperate localities. No species are recorded from Th ailand but we have collected 18 specimens representing 3 or 4 species in Thailand.

Diversity: It is diffi cult to estimate the number of species due to recent changes in generic concepts. Sharkey et al. (2007) divided Coccygidium s.l. into Hypsostypos , Zelomorpha , and Coccygidium s.s., but did not include a list of new combinations so the generic concepts have not been incorporated into the Taxapad database ( Yu et al. 2005). Members of Coccygidium are restricted to the Old World and there are about 10 species recorded for the Oriental region; Bhat and Gupta (1977) included 8 species. Coccygidium corresponds to what they referred to as the Fuliginosa species group of Zelomorpha .

Biology: There are five host records, all on members of the family Noctuidae . Many species are pale colored with rather large ocelli and are nocturnal. The short ovipositor suggests that they are attacking exposed hosts.

Phylogenetic Information. Sister to Zelomorpha which is New World and primarily tropical in distribution ( Sharkey et al. 2006).

Taxonomic Information. Chou and Sharkey (1989) treated Zelomorpha as a junior synonym; however the monophyly of both Coccygidium and Zelomorpha were confirmed by Sharkey et al. (2006).

Diagnosis: The long, style-like, foretibial spur (Fig. 4a) is unique amongst Agathidinae . Members are otherwise very similar to those of Amputostypos , which are more commonly collected in Malaise traps in the Oriental Region.