Deleproctophylla variegata (Klug in Ehrenberg, 1834)

Monnerat, Christian & Abraham, Levente, 2020, Owlflies from Jordan (Neuroptera, Ascalaphidae), Alpine Entomology 4, pp. 139-150 : 139

Deleproctophylla variegata (Klug in Ehrenberg, 1834) Figs 17-23

Ascalaphus variegatus Klug, 1834 - (ODescr), Walker 1953 (Mon), Hagen 1860 (List), 1866 (Tax), McLachlan 1873 (Tax), Navás 1912b (Mon), Alexandrov-Martynov 1926 (Dist), Aspöck et al. 1980 (Mon).

Theleproctophylla barbara auct.(nec Linnaeus) - Navás 1909 (Dist), 1910 (Dist), 1925b (Dist), 1929 (Dist), Bodenheimer, F. S. 1937 (Dist).

Deleproctophylla variegata (Klug, 1834) - Scott 1929 (Dist), Luppova 1973 (Dist), 1966 (Dist), Aspöck et al. 1980 (Mon), Letardi 1991 (Dist), Aspöck and Hölzel 1996 (Chlist), Kačírek 1998 (Dist), Sziráki 2000 (Dist), 2010 (Dist), Háva 2000 (Dist), Koçak and Kemal 2002 (Dist), Whittington 2002 (Dist), Canbulat and Kiyak 2005 (Dist), Canbulat 2007 (Chlist), Dobosz and Ábrahám 2007 (Dist), Krivokhatsky et al. 2015 (Dist).

Material examined.

1♂ Jordania Jordan, Oberes Jordantal, vi.1999, leg. G. Müller, SCMK; 1♂, 1♀, Ayy Alhizman (al-Hazman), 31.11650N, 35.63200E, 1145 m a.s.l., 2.vi.2011, leg. M. Borer, CCM; 2♂♂, NW Wadi Zarqa, 32°11'31.4"N, 36°00'23.2"E, 540 m a.s.l., 8.vi.2011, leg. M. Borer, CCM (Fig. 17 ); 1♀, 1.2 km NE Sakhra, 32,37790N, 35.85935E, 1005 m a.s.l., 9.vi.2011, leg. C. Monnerat, CCM; 2♂♂, 1♀, dir. al-'Aluk, 32.18888N, 35.98066E, 520 m a.s.l., 11.vi.2011, leg. C. Monnerat, CCM; 1♂, 3♀♀, 9.5 km W of Madaba, 31.72070N, 35.69316E, 520 m a.s.l, 26.v.2012, leg. C. Monnerat, CCM (Figs 18 , 22-23 ); 1♂, 3♀♀, 9.5 km W of Madaba, 31.72070N, 35.69316E, 520 m a.s.l, 26.v.2012, leg. M. Borer, CCM (Fig. 21 ); 1♀, 3.vi.2012, Wadi Zarqa, 32.16718N, 36.00473E, 465 m a.s.l., leg. C. Monnerat, CCM; 2♀♀, 3.vi.2012, ar-Rumman, 32.18978N, 35.85004E, 430 m a.s.l., leg. C. Monnerat, CCM; 1♀, 8.vi.2012, between Libb and Muqawir, 31.59344N, 35.68667E, 710 m a.s.l., leg. C. Monnerat, CCM.

Distribution.

The first described Deleproctophylla species was Deleproctophylla variegata (Klug, 1834), known mainly from the West Palaearctic (Europe: Greece (Chios), Asia: Cyprus, Turkey, Caucasus region ( Aspöck et al. 2001), Irak (Kurdistan) ( Scott 1929), Lebanon (type locality), Israel, Turkmenistan, Uzbekistan ( Alexandrov-Martynov 1926), Kyrgyzstan ( Krivokhatsky et al. 2015), Iran (Hamadan; SCMK, unpublished data) and Afghanistan (Upper Silesian Museum, Poland, unpublished data)). It is a new record for the fauna of Jordan. There are false distribution records in Navás’s (1912b) monograph from Spain and France that were corrected in the monograph of Monserrat et al. (2012).

Comments.

It is a day-active species found in open landscapes, for example in extensive cereal fields of Triticum dicoccoides (Figs 19 , 20 ) and fallow fields with Erucaria hispanica . D. variegata is distributed in Jordan in the north and central part of the western highlands area between 430 and 1145 m a.s.l. in semiarid and arid Mediterranean bioclimates. The pterostigma of the specimens collected near Madaba on 26.v.2012 exhibited a gradient of color ranging from beige to light brown to dark brown (Figs 21-23 ), due to maturation processes. Such differences in wing cell coloration were the cause of confusion in the systematics of Deleproctophylla , for example between D. dusmeti ( Navás, 1914) and D. bleusei Kimmins, 1949, as detailed in Monserrat et al. (2014). In mature specimens of D. bleusei , the pterostigma are whitish to bright yellowish, whereas in D. dusmeti the pterostigma are darker. However, other identification criteria such as the wingspot pattern and the morphology of the genitalia are diagnostic and the pterostigma of mature specimens of D. bleusei are whitish to bright yellowish.

Figures

Figures 17 - 24. 17. Habitus of Deleproctophylla variegata male. 18. Habitus of D. variegata female. 19. Habitat of D. variegata, Alhizman, June 2011. 20. Habitat of D. variegata, Wadi Zarqa Valley, June 2011. 21. D. variegata, pterostigma of right forewing, MONNECH 01 _ 004933. 22. D. variegata, pterostigma of right forewing, MONNECH 01 _ 004922. 23. D. variegata, pterostigma of right forewing, MONNECH 01 _ 004921. 24. Habitus of Iranoidricerus cf. iranensis male. Pictures: all C. Monnerat.