"Gulella" (Aenigmigulella) aenigmatica (E. A. Smith, 1890)

Rowson, B, Warren, B. H. & Ngereza, C. F., 2010, Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an " oceanic " island, ZooKeys 70, pp. 1-39 : 19-20

27. "Gulella" (Aenigmigulella) aenigmatica (E. A. Smith, 1890) Figs 57-59

Ennea aenigmatica Smith 1890: 164; pl. 6, fig. 11

Notes.

Pemba material matches Usambara material (NMW) of this Eastern Arc species very well. No species of Aenigmigulella has previously been reported from the coastal region ( Verdcourt 2000, 2006) and it is unknown from Unguja. If native to Pemba it would thus suggest a special Eastern Arc affinity. However we note it was not found in forest reserves apart from a juvenile at Msitu Mkuu FR (Table 2). On Pemba, juvenile shells have complex dentition at a variety of stages. The living animal is cream-coloured. Recent systematic work ( Rowson 2010) indicates "Gulella" aenigmatica does not belong in the genus Gulella L. Pfeiffer and a genus-level revision is in progress.