Harveycapsus dimorpha, Cassis, Gerasimos, Symonds, Celia & Tatarnic, Nikolai, 2010
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publication ID |
https://doi.org/ 10.5281/zenodo.195516 |
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DOI |
https://doi.org/10.5281/zenodo.6206910 |
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persistent identifier |
https://treatment.plazi.org/id/67346D36-FFF9-FE46-DDDC-FB30201CFBA0 |
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treatment provided by |
Plazi |
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scientific name |
Harveycapsus dimorpha |
| status |
sp. nov. |
Harveycapsus dimorpha View in CoL n.sp.
Figures 1–4 View FIGURE 1 View FIGURE 2 View FIGURE 3 View FIGURE 4
Holotype. Male, WESTERN AUSTRALIA: Lochada Nature Reserve, Mungada Ridge, 29°08’11”S 116°54’13”E (WGS 84), 439 m, 18 Sep 2009, M. Harvey, under rocks, banded iron formation (Western Australian Museum; AMNH _PBI0030333). Paratype. Female, same data as holotype (Western Australian Museum; AMNH _PBI0030334).
Diagnosis. This species is recognised by the following combination of characters: male head reddish brown, body medium brown, forewings with pale brown cuneus (Fig, 1a); female body dark brown, almost black, forewings with orange brown markings on margins and along median flexion line and stramineous patch at apex ( Fig. 1 View FIGURE 1 b); frons convex, strongly polished ( Figs 1 View FIGURE 1 a & b); metafemora with three rows of spinules; right paramere club-shaped, denticulate, pubescent ( Fig. 2 View FIGURE 2 b); apex of apophysis of left paramere hooked ( Fig. 2 View FIGURE 2 c); three endosomal spicules ( Fig. 2 View FIGURE 2 f); VES trifurcate ( Fig. 2 View FIGURE 2 f); DES1 bifurcate; DES2 undivided ( Fig. 2 View FIGURE 2 f); and, apex of inter-ramal lobes serrate ( Fig. 3 View FIGURE 3 b).
Description. Male macropterous; female staphylinoid; body length, male 4.51 mm, female 2.78 mm.
Male. COLOURATION. Uniformly medium brown without contrasting markings. Antennae: AI mostly pale brown, AII–AIV medium to dark brown. Head: clypeus, mandibular and maxillary plates dark red; frons medium brown; posterolateral angles of head minor yellowish brown spotting. Labium: LI–LII pale brown; LIII medium brown; LIV dark brown, almost black. Pronotum: anterior margin and midline of callosite region yellow-brown; remainder of callosite region medium brown, a little darker than uniformly medium brown disc. Scutellum : medium brown. Thoracic pleura: proepisternum reddish brown; pterothoracic pleura mostly medium brown; evaporative areas on mesepimeron and metepisternum yellow with red infusion. Forewings: uniformly medium brown, cuneus a little paler, uniformly translucent, membrane fumose, membrane veins pale brown, large membrane cell with small white spot. Legs: coxae and femora yellowish brown, tibiae pale brown, tarsi and pretarsi dark brown. Abdominal venter: medium brown, including terminalia, basally a little paler.
VESTITURE. dorsum with moderate distribution of simple, semi-erect brown setae, more bristlelike on vertex of head and anterolateral angles of pronotum, abdominal venter with simple, semi-erect setae. Antennae: with uniform dense distribution of simple short semi-erect setae, intermixed on AI with handful of elongate dark erect bristlelike setae. Legs: with uniform dense distribution of simple short semi-erect setae; tibiae intermixed with erect bristlelike setae; metafemora with three uniform rows of spinules. Thoracic pleura: uniformly dark brown.
TEXTURE. polished, particularly frons; most of pronotum rugulose, excluding polished callosite region.
STRUCTURE. Antennae: AI elongate, subequal to interocular distance, cylindrical; AII cylindrical, elongate, longer than posterior width of pronotum; AIII little shorter than AII; AIV subequal in length to AI. Labium: greatly elongate, almost reaching pygophore; labial formula L1>LII=LIV>LIII; LI broad, extending well beyond posterior margin of head; LII–LIV bicompressed; LIV strongly tapered. Legs: metafemora ca.
1.6x longer than mesofemora. Male genitalia: right paramere club shaped, unevenly denticulate, densely pubescent ( Fig. 2 View FIGURE 2 b); left paramere with apex of apophysis hooked, sensory lobe thickened, densely pubescent ( Fig. 2 View FIGURE 2 c); VES trifurcate ( Fig. 2 View FIGURE 2 f), deeply dissected, with left hand branch mostly horizontal, with apex serrate and recurved, medial branch most elongate, erect, sinuate, strongly tapered, with dorsal margin serrate, right hand branch ca. 45º to left hand branch, sinuate, strongly tapered, margins serrate; DES1 distally bifurcate, dorsalmost branch serrate; DES2 shortest spicule, sinuate, margins smooth ( Fig. 2 View FIGURE 2 f).
Female. As in male aside from following colouration and vestiture characters. COLOURATION. Mostly dark brown with linear orange brown markings. Head: uniformly dark brown, almost black. Antennae: AI and basal ¾ of AII pale brown, remainder of AII dark brown, base of AIII pale brown, remainder of AIII and AIV dark brown. Labium: female LI pale brown, LII and LIII orange-brown, apex of LIV dark brown. Pronotum: uniformly dark brown, almost black. Scutellum : female dark brown. Forewings: mostly dark brown, with costal margin, narrowly on medium flexion line and claval commissure orange-brown, apex of corium with stramineous marking. Legs: coxae, femora and 4/5th of tibiae pale brown, remainder of tibiae medium brown, first and second tarsomeres pale brown, third tarsomere and pretarsi dark brown, almost black. Abdominal venter: uniformly dark brown. VESTITURE. body with sparse distribution of short, adpressed, pale simple setae, without bristlelike spines, head and pronotum almost glabrous. STRUCTURE. Male genitalia: sclerotized opening of vestibulum produced anteriad with medial margins denticulate ( Fig. 3 View FIGURE 3 a); inter-ramal lobes with apices serrate ( Fig. 3 View FIGURE 3 b).
Distribution. This species is known only from the type locality, in the Avon Wheatbelt IBRA region, southwest Western Australia ( Fig. 4 View FIGURE 4 a).
Ecology. Under rocks on ridgetops in banded iron formation ( Fig. 4 View FIGURE 4 b).
Remarks. The sister-species relationships are unknown. The supraspecific relationships of this taxon are discussed below.
| AMNH |
American Museum of Natural History |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.