Heraeus loja, Dellapé & Melo & Henry, 2016

Dellapé, Pablo M., Melo, María C. & Henry, Thomas J., 2016, A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species, Zoological Journal of the Linnean Society 177 (1), pp. 29-134 : 91

publication ID	https://doi.org/ 10.1111/zoj.12362
publication LSID	lsid:zoobank.org:pub:8A961A76-E33C-4171-B401-0B516B62FA6D
DOI	https://doi.org/10.5281/zenodo.10543739
persistent identifier	https://treatment.plazi.org/id/69250127-E91D-5076-FEA6-3060FAA2E934
treatment provided by	Marcus
scientific name	Heraeus loja
status	sp. nov.

Treatment

HERAEUS LOJA View in CoL SP. NOV.

( FIGS 25I View Figure 25 , 33C View Figure 33 , 34A–D View Figure 34 , 35 View Figure 35 )

Diagnosis

Labium extending to metacoxae. Distiflagellomere with a narrow sub-basal pale band. Membrane brown, with veins paler. Metafemur with a subapical dark band. Heraeus loja sp. nov. shares with H. pallidinervis sp. nov. and H. penai sp. nov. a narrow pale band on the distiflagellomere, occupying at most onequarter of the segment length, and a long labium extending to the metacoxae. These three species, however, are readily distinguished by the male genitalia, with H. loja sp. nov. having a slightly declivent pygophore with narrow, elongate inner projections.

Description ( Fig. 25I View Figure 25 )

Total length 5.89.

Head: Dark brown, shiny; setae long and erect dorsally. Head length 1.20, width 0.93. Postocular length 0.40. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli placed at level of an imaginary line passing the posterior border of eyes. Interocular width 0.48, interocellar width 0.29. Labium pale brown with erect setae, extending to at least metacoxae. Labial segment lengths: I 0.85, II 0.96, III 0.83, and IV 0.35. Antennae pale brown, distiflagellomere with a narrow sub-basal pale band; setae abundant, short, and semierect. Antennal lengths: scape 0.46, pedicel 1.09, basiflagellomere 0.90, and distiflagellomere 0.96. Length of pale band on distiflagellomere 0.13.

Thorax: Anterior pronotal lobe and scutellum dark brown, posterior pronotal lobe paler, with pale spots and with a pale spot on humeral angles. Anterior pronotal lobe with long erect setae, posterior lobe with short erect setae. Collar with scattered punctures, delimited posteriorly by a punctate sulcus. Collar length 0.08, anterior lobe length 0.56, posterior lobe length 0.45; anterior lobe width 1.02, posterior lobe width 1.52. Pleurae dark brown, except acetabular areas reddish brown; punctate, with short recumbent setae. Evaporative area extended. Scutellum dark brown, with a median darker longitudinal stripe, apex paler, pruinose, punctate; with erect setae. Hemelytra mostly pale brown, with irregular darker spots, pattern complex ( Fig. 25I View Figure 25 ). Inner corial spot diffuse; costal margin pale on proximal three-quarters; with a subapical corial spot; membrane brown with veins paler and pale spots between them. Legs: Profemur dark, at most, with apices paler, remainder of legs paler, except metafemur with a subapical dark band ( Fig. 33C View Figure 33 ); setae on profemur long and erect. Protibia curved, protibia and mesofemur without spines.

Abdomen: Dark brown, with abundant short recumbent setae. Male genitalia: Posterior margin of pygophore flattened in dorsal view ( Fig. 34A, B View Figure 34 ), anterior margin of dorsal aperture slightly rounded, with inner projections narrow, elongate, strongly projecting posteriorly. Parameres: Figure 34 View Figure 34 (C, D). Aedeagus: spinose; conjunctiva with spines laterally beneath the ejaculatory reservoir; vesica with sclerotized lobes and spines distally and anteriorly; processus gonopori long and slender.

Distribution

Ecuador ( Fig. 35 View Figure 35 ).

Etymology

The specific epithet is a noun in aposition and refers to Loja Province in Ecuador, where the holotype was collected .

Type material

Holotype: ♂, ECUADOR, Loja, 26 km W of Loja, along road to Catamayo , 2350 m a.s.l., 18-II-2002, 3°59′71″S, 79°18′41″W, T.J. Henry & P.S.F. Ferriera ( USNM).