Kagalana tonde, Bruce, Niel L., 2008

Bruce, Niel L., 2008, New species and a new genus of Cirolanidae (Isopoda: Cymothoida: Crustacea) from groundwater in calcretes in the Pilbarra, northern Western Australia, Zootaxa 1823, pp. 51-64 : 53-57

Kagalana tonde sp. nov.

Material examined (all Pilbarra region, Western Australia).

Holotype. ɗ (2.7 mm) ‘House Creek’, Ashburton, 22°27’52.5”S, 116°02’10.8”E, 8 August 2005, site PSS 172, altitude 150 or 162 m (depending on method of measurement), salinity 0.67 ppm ( WAM C39308).

Paratypes. 2ɗ (2.4, 2.5 [dissected] mm), 3Ψ (non-ovig. 2.8, 2.9, 3.1 mm), same data as holotype ( WAM C39309). 2ɗ (2.2 mm, 1.9 mm), manca (1.5 mm), Hardey River, 22°55’35.2”S, 117°23’03.8” E, 21 November 2003, altitude of 400 m, site code PSS 173, field bore code, NWSLK 220A, coll. Mike Scanlon (MTQ W13692). ɗ (2.6 mm), Hardey River, 22°55’35.2”S, 117°23’03.8” E, 21 November 2003, altitude of 400 m, site code PSS 173, field bore code, NWSLK 220A, coll. Mike Scanlon ( WAM C39311). ɗ (2.7 mm, pereopods 1 both incomplete), 26 July 2003, PSS 173 ( WAM C39310). Ψ (ovig. 3.5 mm), Calgara Bore, 22°58’31.8”S, 116°58’07.3” E, 21 May 2004, altitude of 400 m, site code PSS 209, field bore code, NWSLK 176, coll. Jim Cocking ( WAM C39312). ɗ (2.2 mm), Wyloo Station, 22°39’55.0”S, 116°14’20.2”E, 3 August 2003, altitude of 175 m, site code PSS 206, field bore code, NWSLK 88, coll. Mike Scanlon ( WAM C39313).

Additional material: 5 mancas (1.3–1.6 mm) Robe River 6A, 21°31’53.4”S, 115°512’0.0”E, 11 Nov 2004, PSS 075 ( WAM C39314). Ψ (ovig 3.0 mm; non-ovig, 2.2 mm), Robe River 2A, 21°34’52.6”S, 115°52’7.0”E, 11 Nov 2004, site code PSS 016 ( WAM C39315). 2 mancas (1.8, 1.5 mm), Robe River 2A, 21°34’52.6”S, 115°52’7.0”E, 6 Aug 2005, site code PSS 016 ( WAM C39316). Manca (1.6 mm), Robe River 2A, 21°34’52.6”S, 115°52’7.0”E, 11 Nov 2004, site code PSS 016 ( WAM C39317). Manca (1.6), Robe River 2A, 21°34’52.6”S, 115°52’7.0”E, 15 May 2005, site code PSS 016 ( WAM C39318). Ψ (non-ovig. 2.0 mm), Yarraloola Station, 21°37’58.8”S, 115°57’40.8”E, 5 April 2003, altitude of 70 m, site code PSS 089, field bore code PANNASLK089, coll. Jane Mcrae ( WAM C39319). ɗ (2.0 mm, damaged), 21°37’58.8”S, 115°57’40.8”E, 3 April 2003, site codePSS089, PANNASLK4, coll. Jane Mcrae ( WAM C39320).

Description. Body 2.8 times as long as greatest width, widest at pereonite 6, lateral margins subparallel. Eyes absent. Pereonite 1 and coxae 2–3 each with posteroventral angle rounded; coxae 5–7 with entire oblique carina. Pleon with posterolateral angles of pleonite 2 forming obtuse point; pleonite 3 epimeral margins, narrowly rounded; posterolateral margin of pleonite 4 rounded; pleonite 5 0.86 as wide as anterior margin of pleotelson. Pleotelson 0.9 times as long as anterior width; lateral margins weakly convex, smooth, posterior margin evenly rounded, without median point, with 4 RS.

Antennule peduncle peduncle articles 3 and 4 0.9 times as long as combined lengths of articles 1 and 2, 2.3 times as long as wide; flagellum with 4 articles, extending to anterior of pereonite 1. Antenna peduncle article 4 1.6 times as long as wide, 2.3 times as long as article 3, inferior margin with 1 plumose seta, and 0 short simple setae; article 5 1.3 times as long as article 4, 2.7 times as long as wide (widest sub-distally), inferior margin without pappose setae, superior distal angle with cluster of 2 short simple setae (and 2 plumose setae); flagellum with 9 articles, extending to posterior of pereonite 1.

Frontal lamina 2.1 as long as greatest width, lateral margins diverging slightly towards anterior.

Mandible molar process anterior margin with 12 flat teeth; without proximal cluster of long simple setae; right mandible spine row composed of 4 spines, left with 5 spines; palp article 2 with 7 distolateral setae, palp article 3 with 9 biserrate RS. Maxillule mesial lobe with 3 large CPS RS; lateral lobe with 11 RS. Maxilla lateral lobe with 2 long simple setae (finely serrate); middle lobe with 2 long simple setae (finely serrate); mesial lobe with 2 distal simple setae, with 4 proximal simple and plumose setae. Maxilliped palp article 2 mesial margin with 1 slender seta, lateral margin distally with 1 slender seta (distally serrate); article 3 mesial margin with 4 slender setae, lateral margin with 2 slender setae (distally serrate); article 4 mesial margin with 4 slen- der setae (1 serrate), lateral margin with 1 slender seta; article 5 distal margin 4 setae (2 serrate), lateral margin with 2 setae (1 serrate); endite with 3 long CPS setae, and 1 coupling seta.

Pereopod 1 basis 2.7 times as long as greatest width, superior distal angle with cluster of 2 acute setae; ischium 0.7 times as long as basis, inferior margin without setae, superior distal margin without RS; merus inferior margin with 2 molariform RS, set as two groups, superior distal angle with 2 setae; carpus inferior margin with 1 RS (and 1 long simple seta); propodus 1.7 times as long as wide, inferior margin with 1 RS (spine opposing dactylus with 4 cusps); dactylus 0.7 as long as propodus (slender, 5.0 times as long as basal width). Pereopod 2 ischium inferior margin with 1 stout, bluntly rounded RS, superior distal margin with 1 acute RS; merus inferior margin with 4 stout RS, set as two groups (of 2 and 2), superior distal margin with 2 acute RS; carpus inferodistal angle with 2 RS; propodus 2.5 times as long as wide, palm without RS. Pereopod 3 similar to pereopod 2. Pereopod 6 similar to pereopod 7. Pereopod 7 basis 2.8 times as long as greatest width, superior margin convex, inferior margin with 2 palmate setae; ischium 0.6 as long as basis, inferior margin with 1 RS, superior distal angle with 1 RS (acute), inferior distal angle with 2 RS; merus 0.6 as long as ischium, 1.8 times as long as wide, inferior margin without RS, superior distal angle with 3 RS, inferior distal angle with 1 RS; carpus 0.7 as long as ischium, 1.7 times as long as wide, inferior margin without RS, superior distal angle with 2 RS (1 strongly serrate; plus 1 mid-distal margin), inferior distal angle with 1 RS; propodus 0.9 as long as ischium, 3.4 times as long as wide, inferior margin without RS, superior distal angle with 2 slen- der setae, inferior distal angle with 2 RS. Pereopods 6 and 7, inferior distal angles of ischium to carpus with rounded blunt spines.

Pleopod 1 exopod 1.5 times as long as wide, distally broadly rounded, lateral margin weakly convex, mesial margin weakly convex, with PMS from distal one-third, with 9 PMS; endopod distally subtruncate, lateral margin straight, with 4 PMS on distal margin only; peduncle 1.4 times as wide as long, mesial margin with 3 coupling hooks. Pleopod 2 exopod with 10 PMS, endopod with 1 PMS; appendix masculina with parallel margins, 1.2 times as long as endopod, distally bluntly rounded. Pleopod 3 exopod with 8 PMS. Pleopod 4 exopod with 10 PMS. Pleopod 5 exopod with 12 PMS. Pleopods 2–5 peduncle distolateral margin with prominent acute RS, 3–5 endopods without distomesial serrate scales.

Uropod peduncle ventrolateral margin with 2 RS; lateral margin without medial short acute RS, uropod peduncle posterior lobe about three-quarters as long as endopod. Uropod rami extending to pleotelson apex, marginal setae in single tier, apices narrowly rounded. Endopod apically not bifid; lateral margin sinuate, proximal lateral margin without RS; distal lateral margin with 1 RS, mesial margin straight (serrate), with 3 RS. Exopod not extending to end of endopod, 3.2 times as long as greatest width, apically not bifid; lateral margin weakly convex, without RS; mesial margin weakly convex, with 1–3 RS.

Variation. Details of type specimens. Pleotelson (n =13) 2–4 RS, with 4 most frequent (64%), 2 occurring three times and 3 twice. Uropod endopod (n = 25) mesial margin with 2–5 RS, with 2 (40%) and 3 (56%) most frequent (5 once), lateral margin (n = 23) with 0 (39%) or 1 (61%) RS; exopod mesial margin (n = 25) with 1– 3 RS, with 1 (40%), or 2 (57%) most frequent (3 once), lateral margin (n = 25) with 1 (88%) or 2 (12%) RS.

Etymology. Tonde is an Aboriginal word meaning blind (noun in apposition).