Pygatyphella ( Ballantyne 1968 )

Ho, - Z., 2019, The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species, Zootaxa 4687 (1), pp. 1-174 : 119

Pygatyphella ( Ballantyne 1968)

Figs 30−37

Luciola (Pygatyphella) Ballantyne 1968: 109 .

Pygatyphella (Ballantyne) . Ballantyne & Lambkin 2009: 77 , figs 23–26, 61, 69–72, 255–328, 377–396, 441–487.

Type species: Atyphella obsoleta Olivier 1911b .

Diagnosis. Both Pygatyphella and Pacifica occur in New Guinea and Pacific islands, and both have LO in V7 retracted to the anterior half or less; the posterior clear portion of V7 is frequently arched and underlying muscle impressions can be clearly seen. The two genera are most obviously distinguished by their dorsal colouration, Pygatyphella having a pattern resembling bird droppings, where most Pacifica species have orange pronotum and dark brown elytra which may be pale margined. Ballantyne & Lambkin (2009) distinguished two forms within Pygatyphella they named Pygatyphella A and B. Subsequently Pygatyphella B was described from 5 species as a distinct genus Pacifica . The following diagnosis of Pygatyphella sensu Ballantyne & Lambkin (2013) is modified from Ballantyne & Lambkin (2009): Dorsal colouration usually cryptic, very pale brown with dark markings on pronotum, MS, MN and elytral base and apex; pronotum often with angulate convergence along lateral margins, and small depression often present just anterior to posterolateral corner; lacking irregularities at the corners; posterolateral corners may project beyond median posterior margin; at least two interstitial lines usually well–defined; elytral margins convex or parallel-sided; posterior area of V7 may reach into LO (which is medially emarginated); apex of MPP rounded, pointed, or squarely or obliquely truncate; MPP lacking median dorsal ridge (except in Pygat. karimui), lacking curved, slightly off-centre posterior ridge on ventral surface of T8; dorsal surface of posterior area of MPP faced with cuticle that is not attached to the ventral surface of V7 and ends just behind the area of muscle attachment posterior to the LO; the anterior margins of this cuticle are reflexed in Pygat. peculiaris; median area of this facing cuticle lacking longitudinal split; T8 anterolateral prolongations lacking ventral projections at their base; aedeagal sheath not subparallel-sided in basal third; aedeagus L/W usually>3; LL/ML not wide; ML sometimes asymmetrical in a horizontal plane only, never with lateral teeth; anterior margin of LL often asymmetrically produced.

Figures

FIGURES 30−37. Key to genera characters 5. 30, 34, 35 Pygatypella undulata; 31 Pygat. wisselmerenia; 32, 36 Pygat. obsoleta; 33 Pygat. peculiaris; 37 Pygat. kiunga paratype male. 30−34, 37 dorsal (34 diagrammatic representation of colour pattern); 35 aedeagal sheath, ventral; 36 aedeagus ventral, line outline.