Caraphia granulifera Holzschuh, 1984

Ohbayashi, Nobuo, Lin, Mei-Ying & Yamasako, Junsuke, 2016, Revision of the Caraphiini, New Tribe (Coleoptera, Cerambycidae, Lepturinae), Zootaxa 4084 (2), pp. 187-217 : 191-192

2. Caraphia granulifera Holzschuh, 1984

( Figs 9, 10, 11 , 44 , 51 , 67 , 83 , 99 , 115 , 129 )

Caraphia granulifera Holzschuh, 1984: 141 , fig. 1 (Type locality: Dhunche, Trisuli Khola, Nawakot, C-Nepal); Weigei, 2006: 497; Chou & N. Ohbayashi, 2008: 141.

Diagnosis. Male: BL=13.0– 14.3 mm; EW= 3.6–3.8 mm. Body mostly reddish brown. Head, pronotum and scape with dense yellowish scales. Head with inter-ocular distance very wide, nearly half of head width; antennae slightly exceeding elytral apex; relative lengths of segments from base to apex: 50: 15: 54: 54: 80: 78: 74: 71: 75: 69: 80.

Pronotum distinctly wider than long, constricted at apical and basal fourth; sides triangularly inflated laterally; disk with a pair of small protuberant behind anterior constriction.

Elytra narrowed from base to apex, foveate in regular rows except for along suture and apical area which are irregularly arranged; scales pale yellow, relatively long and thin, suberect and barely curved, 2.0 times as long as the diameter of foveae, mostly alternating on rows of foveae.

Genitalia ( Fig. 115 ) with paramere of tegmen wide and short, 2.44 times as long as basal width, slightly attenuate toward rounded apices, 0.17 times as long as tegmen. Median lobe with dorsal plate almost reaching the apex of ventral plate; basal struts slightly shorter than a half of median lobe. Endophallus ( Fig. 129 ) rather simple and elongate, MPH roundly inflated and with indistinct sclerites near base; APH claviform with a pair of small dorsal projections near base, without rod-like sclerite.

Female: BL= 14.1 mm; EW= 3.8 mm. Similar to male in body color, foveae and scales, but antennae distinctly short, reaching apical fifth of elytra; relative lengths of segments from base to apex: 47: 14: 49: 46: 65: 58: 55: 51: 51: 49: 60. Elytra slightly widened from humeri to apical fourth.

Type materials examined. Holotype: ♂, C-Nepal, Nawakot, Trisuli Khola , 2,200 m, Dhunche , 15–21-X- 1982, C. Holzschuh leg., Geschlüpft aus Dürren Ästen: Quercus Imago in Puppenviege ( PCCH). Paratype: 1♀, same data as holotype, Geschlüpft aus Dürren Ästen: IX. 1982 ( PCCH).

Other material examined. 1♂, same data as holotype, Geschlüpft aus Dürren Ästen: Rhododendron , 6-II- 1986 ( PCCH).

Distribution. Nepal.

Remarks. This species is most similar to C. cribrata , but easily distinguishable by its short antennae, wide inter-ocular distance, and indistinct elytral carinae. According to the original description, the adults were reared from Quercus sp. and Rhododendron sp.

Figures

FIGURES 6 – 22. Habitus and type-labels of Caraphia spp. 6, 7, 8. Caraphia cribrata (6. ♂; 7. holotype, ♀; 8. ditto, type label); 9, 10, 11. C. granulifera (9. holotype, ♂; 10. ditto, label; 11. paratype, ♀); 12, 13. C. minor (12. lectotype ♀; 13. ditto, label); 14, 15, 16. C. depressa (14. holotype, ♂; 15. ditto, label; 16. paratype, ♀); 17, 18. C. ebenina (17. ♂; 18. ♀); 19, 20. C. laosica (19. holotype, ♂; 20. ditto, label).

FIGURES 37 – 49. Habitus, type-labels and hind wing of Caraphia spp. 37, 38. Caraphia paulastoni sp. nov. (37. holotype ♂; 38. paratype ♀); 39. C. squamosa comb. nov., ♀; 40, 41. C. seriata comb. nov. (40. holotype, ♀; 41. ditto, label); 42. C. lingafelteri sp. nov. (holotype, ♂); 43. C. cribrata; 44. C. granulifera; 45. C. depressa; 46. C. ebenina; 47. C. borneana; 48. C. squamosa, comb. nov.; 49. C. lingafelteri, sp. nov.

FIGURES 50 – 65. SEM images (400 – 500 ×) of pronotal sculpture and scales of Caraphia spp. 50. Caraphia cribrata; 51. C. granulifera; 52. C. minor; 53. C. depressa; 54. C. ebenina; 55. C. laosica; 56. C. thailandica; 57. C. borneana; 58. C. huai sp. nov.; 59. C. laticeps (Pic, 1922); 60. C. lepturoides lepturoides; 61. C. taiwana; 62. C. paulastoni sp. nov.; 63. C. squamosa, comb. nov.; 64. C. seriata, comb. nov.; 65. C. lingafelteri, sp. nov.

FIGURES 66 – 81. SEM images (120 – 150 ×) of elytral foveae and scales of Caraphia spp. 66. Caraphia cribrata; 67. C. granulifera; 68. C. minor; 69. C. depressa; 70. C. ebenina; 71. C. laosica; 72. C. thailandica; 73. C. borneana; 74. C. huai sp. nov.; 75. C. laticeps; 76. C. lepturoides lepturoides; 77. C. taiwana; 78. C. paulastoni sp. nov.; 79. C. squamosa, comb. nov.; 80. C. seriata, comb. nov.; 81. C. lingafelteri, sp. nov.

FIGURES 82 – 97. SEM images (400 – 500 ×) of elytral foveae and scales of Caraphia spp. 82. Caraphia cribrata; 83. C. granulifera; 84. C. minor; 85. C. depressa; 86. C. ebenina; 87. C. laosica; 88. C. thailandica; 89. C. borneana; 90. C. huai sp. nov.; 91. C. laticeps; 92. C. lepturoides lepturoides; 93. C. taiwana; 94. C. paulastoni sp. nov.; 95. C. squamosa, comb. nov.; 96. C. seriata, comb. nov.; 97. C. lingafelteri, sp. nov.

FIGURES 98 – 113. Alignment of scales of Caraphia spp. near middle of elytra. 98. Caraphia cribrata; 99. C. granulifera; 100. C. minor; 101. C. depressa; 102. C. ebenina; 103. C. laosica; 104. C. thailandica; 105. C. borneana; 106. C. huai sp. nov.; 107. C. laticeps; 108. C. lepturoides lepturoides; 109. C. taiwana; 110. C. paulastoni sp. nov.; 111. C. squamosa, comb. nov.; 112. C. seriata, comb. nov.; 113. C. lingafelteri, sp. nov.

FIGURES 114 – 121. Male genitalia of Caraphia spp. 114. Caraphia cribrata; 115. C. granulifera; 116. C. depressa; 117. C. ebenina; 118. C. laosica; 119. C. thailandica; 120. C. borneana; 121. C. huai sp. nov.

FIGURES 128 – 136. Endophallus of Caraphia spp. 128. Caraphia cribrata; 129. C. granulifera; 130. C. depressa; 131. C. ebenina; 132. C. laosica; 133. C. thailandica; 134. C. huai sp. nov.; 135. C. laticeps (Pic, 1922); 136. C. lepturoides lepturoides; 137. C. taiwana; 138. C. paulastoni sp. nov.; 139. C. lingafelteri, sp. nov.