Tomarus paranaensis, Lopez-Garcia & Deloya, 2019

López-García, Margarita M. & Deloya, Cuauhtémoc, 2019, Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species, The Coleopterists Bulletin 73 (1), pp. 127-141 : 133

Tomarus paranaensis López-Garc´ ıa and Deloya, new species

Zoobank.org/ urn:lsid:zoobank.org:act:5FF67FC3-A050-475F-9541-D42E2CCFFA26 ( Figs. 17–19 )

Type Material. 16. Holotype labeled “ Ponta Grossa / Paraná / 3.ii 1965 /Moses / 26438 // Ligyrus gibbosus DeGeer // Halik det. 1965 // BRASIL Halik / 1966 / Collection” (16 USNM).

Description. Holotype. Habitus as in Fig. 17 . Length 15.5 mm; width across humeri 8.7 mm. Color reddish brown. Head: Frons coarsely punctate, area between eyes nearly impunctate. Frontoclypeal region with 2 cariniform tubercles separated by less than 1 tubercle diameter. Clypeus slightly narrowed towards apex, base 2 times as wide as apex. Clypeal teeth triangular, separated by a tooth diameter. Mandibles with 2 apical, acute teeth and a widely rounded basal lobe. Interocular distance equal to 5 times eye width. Antennal club long, 1.5 times longer than antennomeres 2–7. Pronotum: Surface with small, dense punctures. Apical tubercle and fovea absent. Margin of apical angles entire. Elytra: First interval punctate, punctures similar to those on other intervals. Pygidium: Surface with small, evenly distributed punctures; not rugose Punctures small, evenly distributed. Strongly convex in lateral view. Apex regularly rounded. Legs: Protibia tridentate without an additional small basal tooth. Protarsus subcylindrical, not enlarged; inner claw entire. Metatibia strongly narrowed before apex. Apex of metatibia entire, with 19 spinules. Parameres: With a medial tooth on each side; apex widened but ending in an acute, triangular point ( Figs. 18–19 ).

Etymology. The species’ name refers to the type locality in the state of Paraná, Brazil.

Distribution. The holotype was collected in Ponta Grossa ( Fig. 23 ), a municipality located in the Second Plateau of the state of Paraná ( Brazil) within the Atlantic Forest biome, represented by the Araucaria Forest ecosystem ( Mielke et al. 2012).

Temporal Distribution. February (1).

Diagnosis. Tomarus paranaensis and T. burmeisteri are similar in their external morphology. However, the metatibiae are strongly narrowed before the apex and have 19 spinules on the apical margin in T. paranaensis , whereas they have the sides nearly parallel and the apical margin with 25–27 spinules in T. burmeisteri . The parameres are apically acute in T. paranaensis ( Figs. 18–19 ) but rounded to subquadrate in T. burmeisteri (Fig. 49; see also fig. 18 in Neita-Moreno and Ratcliffe 2017).

Figures

Figs. 17–19. Tomarus paranaensis. 17) Habitus; 18) Parameres, frontal view; 19) Parameres, lateral view. Scale bars = 2 mm (Fig. 17), 0.5 mm (Figs. 18–19).

Fig. 23. Known localities of five newly described species of Tomarus from Central and South America.