Ischnopodaumbratica ( Erichson, 1837 ) PA–NIK, 2006

PA – NIK, GRZEGORZ, 2006, Taxonomy and phylogeny of the World species of the genus Ischnopoda Stephens, 1837 (Coleoptera, Staphylinidae: Aleocharinae), Zootaxa 1179 (1), pp. 1-96 : 27-34

Ischnopodaumbratica ( Erichson, 1837) , comb. n. ( Figs. 19–21 )

Tachyusa umbratica Erichson, 1837: 310 .

Tachyusa umbratica: Heer, 1839: 345 .

Tachyusa (Ischnopoda) umbratica: Ganglbauer, 1895: 245 .

Ischnopodaumbratica: Reitter, 1909: 73 .

Tachyusa (Ischnopoda) umbratica: Bernhauer & Scheerpeltz, 1926: 584 .

Tachyusa (Ischnopoda) umbratica: Lohse, 1974: 70 .

Type material. Lectotype (here designated): ♂: Hist.­Col. ( Coleoptera ) Nr. 5338, Tachyusa umbratica Erichson , Austria, Ullr. ( ZMHB). Paralectotypes: 2♂♂ and ♀: same data as the lectotype ( ZMHB).

Additional material. Albania: Krujë , Mader, 2 exx ( NMW) . Austria: Albern , Donauauen, Scheerpeltz, 5 exx ( MHNG) ; 68 exx ( NMW); Albern near Wien, 21.IV.1918, Curti, 2 exx ( NMW) ; Bisamberg , 2 exx ( NMW) ; Burgenland, Neusiedlersee, Leitha­ Aatarm , 26.V.1985, 3 exx ( VAPC) ; Eisenkappel, Karten , VII–VIII.1926, Scheerpeltz, 4 exx ( NMW) ; VII–VIII.1931, Scheerpeltz, 13 exx ( NMW) ; 7.VIII.1931, Scheerpeltz, 1 ex ( NMW) ; Enzersdorf , Wagner, 1 ex ( NMW) ; Enzersfeld , Luze, 2 exx ( NMW) ; Graz , Messa, 2 exx ( MSNM) ; Graz environs, Netolitzky, 1 ex ( NMW) ; Hochschwab, Bodenbauer environs, ca. 850–1000 m, 26.VII–5.VIII.1921, Scheerpeltz, 1 ex ( NMW) ; Horn , Bernhauer, 1 ex ( FMNH) ; Inzeradorf near Wien, Lieeingbach river , 28.V.1946, P. Meyer, 1 ex ( NMW) ; Kainach river, Lannach St. , 3 exx ( MIZ) ; Karawanken, Draubanke bei Ferlach , 1–12.VIII.1924, Scheerpeltz, 4 exx ( NMW) ; Klosterneuburg, Donau Auen , 18.IV.1946, P. Meyer, 4 exx ( NMW) ; Krems , Th. Wanka, 1 ex ( CNC) ; 2 exx ( FMNH); Leopoldsdorf near Wien, 17.VI.1932, Netolitzky, 2 exx ( NMW) ; Linz, H. Priesner, 5 exx ( NMW) ; Modling near Wien, 26.V.1942, Scheerpeltz, 36 exx ( NMW) ; Neudorf , 1 ex ( NMW) ; Oberlaa near Wien, 23.IV.1949, P. Meyer, 3 exx ( NMW) ; Plankenberg , 1 ex ( NMW) ; Radstadtar Tauer , Schladming, 1 ex ( NMW) ; Rodaun, Kaltendeuteeben , 12.VI.1942, Netolitzky, 1 ex ( NMW) ; Sankt Aegyd am Neuwalde , 1 ex ( NMW) ; Seremering , Lang, 1 ex ( NMW) ; Staubeckew near Wien, 26.IV.1946, P. Meyer, 1 ex ( NMW) ; Steinegg , 2 exx ( FMNH) ; Steyr environs, Petz, 1 ex ( NMW) ; Stockerau , Bernhauer, 1 ex ( FMNH) ; Luze , 2 exx ( NMW) ; Skalitzky , 1 ex ( NMW) ; Wassergefullte, Bombeutrichter near Inzersdorf near Wien, 24.IV.1946, P. Meyer, 1 ex ( NMW) ; Weisskirchen, V. Zoufal, 2 exx ( NMW) ; Wien, Donauauen, 3 exx ( NMW) ; Wien, Breit, 2 exx ( NMW) ; Curti , 5 exx ( NMW) ; Moczarski , 1 ex ( NMW) ; Skalitzky , 1 ex ( MNHN) ; Reitter , 1 ex ( ISEA) ; 1 ex ( MHNG); 1 ex ( MNHN); 1 ex ( NHMB); Wien, Preter , 8.VI.1946, P. Meyer, 1 ex ( MHNG) ; Wien, Tullnerbach, 1 ex ( ISEA) ; Wienerwald , Breit, 4 exx ( NMW) . Azerbaijan: Lenkoran, Leder, 1 ex ( HNHM) . Belgium: Anderlecht, Pede valley , 16.IV.1945, Fagel, 3 exx ( IRSNB) ; Angre bois d'Angre , 12.VI.1947, Fagel, 7 exx ( IRSNB) ; Anseremme la Lesse , 31.V.1946, Fagel, 1 ex ( IRSNB) ; Anvers , 11.V.1938, Fagel, 1 ex ( IRSNB) ; Ben­Ahin , 9.VI.1941, Fagel, 1 ex ( IRSNB) ; Bolderberg , 7.VII.1953, Fagel, 6 exx ( IRSNB) ; Evrehailles, Bauche , 17.V.1947, Fagel, 4 exx ( IRSNB) ; Hastière­ Lavaux Maurenne , 29.VIII.1946, Fagel, 1 ex ( IRSNB) ; Ittrebeek, Pede valley , 9.V.1945, Fagel, 3 exx ( IRSNB) ; Nameche , La Meuse, 4.VI.1939, Fagel, 1 ex ( IRSNB) ; Nismes , 16.V.1948, Fagel, 2 exx ( IRSNB) ; Ruysbroeck , 17.IV.1943, Fagel, 2 exx ( IRSNB) ; Seilles, Sclaigneaux , 17.VIII.1945, Fagel, 2 exx ( IRSNB) ; Seilles , 15–20.VI.1943, Fagel, 3 exx ( IRSNB) ; Thon, Samson , 4.VI.1939, Fagel, 1 ex ( IRSNB) ; Vieuxville, Ourthe river , 25.IV.1953, Fagel, 9 exx ( IRSNB) ; Vilvorde , 26.VII.1942, Fagel, 2 exx ( IRSNB) ; Walzin , 31.V.1946, Fagel, 2 exx ( IRSNB) ; Wemmel , 9.V.1945, Fagel, 4 exx ( IRSNB) . Bosnia and Herzegovina: Barnik , 23.V.1972, 1 ex ( MHNG) ; Buna , 13.VII.1977, 1 ex ( MHNG) ; Ilid _a, Apfelbeck, 2 exx ( FMNH) ; K. Kysely, 1 ex ( MHNG); 9.VI.1911, 1 ex ( NMPC) ; Prozor , 1902, O. Leonhard, 1 ex ( NMW) . Bulgaria: Melnik , 14.VII.1983, Wrase & Schülke, 3 exx ( ZMHB) ; Rupite near General Todorow , 23–28.IV.1984, Hieke, 1 ex ( ZMHB) ; Sandanski­Struma , 28.IV.1985, Jaeger, 2 exx ( ZMHB) ; Schwarzmeerkuste, Vlas near Hessebar , 7–15.V.1985, Wrase, 1 ex ( ZMHB) ; Sofia, 1908, Rambousek, 3 exx ( NMPC) ; Vles near Nesebar , 17–26.VII.1983, Wrase & Schülke, 6 exx ( ZMHB) . Croatia: Fuzine , Bernhauer, 1 ex ( FMNH) ; Zagreb, VII.1955, Coiffait, 1 ex ( MNHN) ; V. Stiller , 27.VI.1917, 1 ex ( HNHM) . Czech Republic: “Brandeis" [Brandys], Skalitzky, 2 exx ( NMW) ; 11.V.1893, 1 ex ( ZFMK) ; 7.V.1894, 1 ex ( MSNM) ; Branik , 16.IV.1906, Rambousek, 1 ex ( NMPC) ; Zeman , 2 exx ( NMPC) ; 11.V.1892, Missionshaus, 1 ex ( ZFMK) ; Brod, T. Wanka, 1 ex ( NMW) ; Carlsbad [Karlovy Vary], Rodt, 12 exx ( NMPC) ; 1 ex ( NMW); Čejč u Hod, J. Bechyne, 3 exx ( NMPC) ; Čelakovice , 7.VI.1909, Klička, 2 exx ( NMPC) ; 19.IV.1917, 2 exx ( NMPC) ; 1923, Rambousek, 1 ex ( NMPC); 17.V.1960, 2 exx ( MHNG) ; Rambousek , 1 ex ( NMPC) ; Heyrovsky , 1 ex ( NMPC) ; Klička , 2 exx ( NMPC) ; České Sredo near Litomėřice , 23.V.1983, Wrase, 3 exx ( ZMHB) ; Chrudim , Ručicka, 1 ex ( NMPC) ; Chuchel , Zeman, 1 ex ( NMPC) ; Cibulka , 5.V.1906, Rambousek, 1 ex ( NMPC) ; Davle , Zeman, 1 ex ( CAS) ; 1 ex ( NMPC); 4 exx ( NMW); Dobřichovice , 4.V.1906, 2 exx ( NMPC) ; Dobřiš , Jelinek, 2 exx ( NMPC) ; Hlubočepy , Jouki, 3 exx ( NMPC) ; Jilovištì , 24.IV.1910, 1 ex ( NMPC) ; “ Jungbunzlau " [Mladá Boleslav], Skalitzky, 6 exx ( NMW) ; Kárany , 29.V.1910, 1 ex ( NMPC) ; Kic , Klička, 7 exx ( NMPC) ; Klànovice, K. Pfleger, 1 ex ( NMPC) ; Kováčová , VI.1935, 2 exx ( NMPC) ; Lety , 19.IX.1909, Heyrovsky, 2 exx ( NMPC) ; Libice , 26.VI.1907, Klièka, 2 exx ( NMPC) ; Lobkovice , Zeman, 3 exx ( NMPC) ; Malá Skála , Obenberger, 2 ex ( NMPC) ; Moravičany , 7.V.1970, O. Marek, 1 ex ( FMNH) ; Mutĕnice, J. Bechyne, 1 ex ( NMPC) ; Neratovice , Heyrovsky, 2 exx ( NMPC) ; 29.IX.1918, 5 exx ( NMPC) ; Oskau , Bernhauer, 3 exx ( FMNH) ; Pardubice, Zeman, 1 ex ( NMPC) ; “ Paskau " [Paskov], Graf, 2 exx ( MIZ) ; Reitter , 3 exx ( HNHM) ; Reitter , 1 ex ( MCZ) ; Reitter , 1 ex ( MHNG) ; Reitter , 1 ex ( NMPC) ; Pisty , 8.IX.1907, Klička, 2 exx ( NMPC) ; Plasy , Heyrovsky, 1 ex ( NMPC) ; Plžen, Kouril, 3 exx ( NMPC) ; Přibram , 6 exx ( NMPC) ; “ Prossnitz " [Prostejov], V. Zoufal, 2 exx ( MIZ) ; 12 exx ( NMW); 1 ex ( MTD); Štĕchovice , Zeman, 1 ex ( CAS) ; 3 exx ( NMPC); Dvorak , 1 ex ( CNC) ; Stromovka , 10.VII.1910, Kracik, 1 ex ( NMPC) ; Tišice, O. Kavan, 1 ex ( NMPC) ; Toušeń , 25.IV.1903, 25 exx ( NMPC) ; Heyrovsky , 1 ex ( NMPC) ; Vrané , 21.VI.1909, Jedlicka, 1 ex ( NMPC) ; 21.V.1911, 2 exx ( NMPC) ; Závist , 1.V.1910, Obenberger, 1 ex ( NMPC) ; Zlechov , 11.VI.1905, Rambousek, 1 ex ( NMPC) ; 19.III.1906, 2 exx ( NMPC) ; 1.VII.1919, 3 exx ( NMPC) . Denmark: Kopenhagen, V. Hansen, 1 ex ( MHNG) . France: Achères , VII.1912, 6 exx ( BMNH) ; Alpes Maritimes, Lombat , 11.IV.1933, Torre Tasso, 1 ex ( MSNM) ; Alsace, 2 exx ( CAS) ; Ancenis , 4 exx ( MNHN) ; Anjou , 3 exx ( MNHN) ; Aveyron near Penne, Tarn , 27.VI.1976, 1 ex ( MHNG) ; Bareges , 12.VIII.1970, Roegemont, 1 ex ( GRPC) ; Bonneuil , VIII.1938, Levasseur, 1 ex ( MNHN) ; Briare , VII.1942, 1 ex ( MNHN) ; V.1943, 2 exx ( MNHN) ; Brignais , Rhone, G. Audras, 1 ex ( MHNL) ; Brivé , V.1944, Coiffait, 1 ex ( MNHN) ; Chabanais , VIII.1940, 1 ex ( MNHN) ; Charentais , Bord du l'Armencon, 30.IV.1938, 4 exx ( MNHN) ; Chartreuse , 1 ex ( BMNH) ; Chateauroux , V.1938, Coiffait, 1 ex ( MNHN) ; Chatou , V.1936, 1 ex ( MNHN) ; VI.1939, 5 exx ( MNHN) ; Contes, Alpes­Martimes , Deville, 2 exx ( MNHN) ; Dourbie, Millau , V.1956, 2 exx ( MHNG) ; Écully , Rhone, G. Audras, 7 exx ( MHNL) ; Gevrieux, G. Audras, 4 exx ( MHNL) ; Hyeres , 2 exx ( MNHN) ; Jemelle , La Lomme, 18.VI.1939, Fagel, 5 exx ( IRSNB) ; L'Escarene, Alpes­Maritimes , Deville, 1 ex ( MNHN) ; Le Legue, 9.VII.1938, Levasseur, 1 ex ( MNHN) ; Le Buc , V.1905, 6 exx ( BMNH) ; Lent, G. Audras, 1 ex ( MHNL) ; Lent, G. Audras, 1 ex ( MHNL) ; Les Sables d'Olonne, G. Audras, 1 ex ( MHNL) ; Lompret , 26.VIII.1962, 1 ex ( IRSNB) ; Marly , Levasseur, 1 ex ( MNHN) ; Marseille , 1 ex ( MNHN) ; Morzine environs near Le Lindatet, 1400 m, 12.VIII.1977, Puthz, 1 ex ( MHNG) ; Mulhouse­Ill, Mount Rhin , VI.1952, O. Schmitt, 4 exx ( MNHN) ; Paris , 7 exx ( MNHN) ; Remilly, Moselle , VIII.1922, 1 ex ( MNHN) ; Riofreddo , Roffiny, 2 exx ( MNHN) ; Ruisseau, St. Leonard , 27.VII.1927, 1 ex ( MNHN) ; Saint Germain , Rhône, G. Audras, 1 ex ( MHNL) ; Saint Dizier , Marne, Deville, 3 exx ( MNHN) ; Saint Loup , 27 exx ( MNHN) ; Saint Martin , VI.1951, Levasseur, 1 ex ( MNHN) ; Saint Valery , 2 exx ( BMNH) ; Saint Rambert, G. Audras, 2 exx ( MHNG) ; Saint Florent , Cher, Deville, 3 exx ( MNHN) ; Soissons , 20.V.1905, 1 ex ( MNHN) ; Strasbourg , VII.1952, 1 ex ( MHNG) ; 9.V.1953, Coiffait, 3 exx ( MNHN) ; Triel , IV.1903, 1 ex ( BMNH) ; Vald Isere, G. Audras, 1 ex ( MHNL) ; Vaucluse, Cecile , V.1945, Coiffait, 1 ex ( MNHN) ; 6.VII.1969, 4 exx ( BMNH) ; Villeneuve , VI.1947, Levasseur, 1 ex ( MNHN) ; Vitry , VII.1938, 1 ex ( MNHN) ; Wassy, Marne , Deville, 1 ex ( MNHN) ; Wissous , V.1938, 29 exx ( MNHN) . Germany: Aussig, 1 ex ( MHNG) ; 2 km SW Wildenthal , 12.V.1988, Uhlig, 1 ex ( ZMHB) ; Barnitz , 4.V.1880, 2 exx ( MHNG) ; Bayern, Assernaver , 11.VI.1932, Ihssen, 1 ex ( ZMHB) ; Bayern, Birach near München , 22.V.1931, Ihssen, 2 exx ( ZMHB) ; Bayern, Garmisch, Loisach river , 4.VII.1932, Ihssen, 1 ex ( ZMHB) ; Bayern, Unterbruck , 8.VI.1931, Ihssen, 1 ex ( ZMHB) ; Berlin, Brieselang, 8 exx ( DEI) ; Berlin, Friedrichshagen , 10–12.VII.1981, U. Gollner, 1 ex ( ZMHB) ; 30.VI.1987, U. Gollner, 2 exx ( ZMHB) ; 7.VII.1987, U. Gollner, 2 exx ( ZMHB) ; 10–11.VII.1987, U. Gollner, 2 exx ( ZMHB) ; 21–22.VII.1987, Gollner, 1 ex ( ZMHB) ; Berlin­Biesdorf , 15.VIII.1979, H. Wendt, 1 ex ( ZMHB) ; 24–26.VII.1982, U. Gollner, 1 ex ( ZMHB) ; 25.VII.1982, K. Gunther, 3 exx ( ZMHB) ; 4.VIII.1982, U. Gollner, 1 ex ( ZMHB) ; 15–16.VII.1983, U. Gollner, 1 ex ( ZMHB) ; Beuern near Giessen , 5.V.1949, Banninger, 6 exx ( NMW) ; Borussia , 1 ex ( CAS) ; Buckow Schermutzelsee Sandufer , 10.V.1981, Uhlig, 1 ex ( ZMHB) ; Carlshafen , 1.V.1934, Folwaczny, 1 ex ( MHNG) ; Chemnitz , Kirsch, 3 exx ( MTD) ; Dresden , 13.IV.1914, H. Koksch, 1 ex ( MTD) ; 1 ex ( FMNH); Eberswalde environs, Werbellinsee , SW Bucht, 24.IV.1983, Uhlig, 1 ex ( ZMHB) ; Eilenburg, Prov. Sachsen, 6.VII.1958, Linke, 1 ex ( MTD) ; Erfurt , 17 exx ( NMW) ; Geesthacht near Hamburg, 3.VI.1977, Koltze, 1 ex ( DEI) ; Golm environs, 1 ex ( DEI) ; Halnburger Berge , Scheerpeltz, 5 exx ( NMW) ; Hanau a. Main , V.1911, 1 ex ( FMNH) ; Hannover , 29.VII.1989, 5 exx ( VAPC) ; Leine river , 6.V.1990, 3 exx ( VAPC) ; 11.IV.1991, Assing, 1 ex ( VAPC) ; Herzfelde near Rudersdorf, 3 exx ( DEI) ; Kaitz Gr. , V.1923, K. Hanel, 3 exx ( MTD) ; Kleinheringen near Bad Kosen , 8.V.1986, F. Hieke, 1 ex ( ZMHB) ; Konigsberg , 5 exx ( NMW) ; Krumme Hufe, Naumburg , 12.V.1922, Maertens, 3 exx ( ZMHB) ; Leipzig , 29.IV.1905, Linke, 1 ex ( MTD) ; 4.V.1907, 2 exx ( MTD) ; 29.VI.1907, 1 ex ( MTD) ; 18.VII.1907, 1 ex ( MTD) ; 4.VIII.1907, 9 exx ( MTD) ; 29.IX.1907, 1 ex ( MTD) ; 12.IX.1909, 1 ex ( MTD) ; 9.VI.1926, 1 ex ( MTD) ; 12.V.1929, 2 exx ( MTD) ; 12.VIII.1934, 1 ex ( MTD) ; 14.IX.1934, 1 ex ( MTD) ; 14.V.1935, 1 ex ( MTD) ; 2.V.1937, 5 exx ( MTD) ; 10.V.1942, 2 exx ( MTD) ; 23.VII.1944, 1 ex ( MTD) ; 10.VI.1961, K. Dorn, 1 ex ( ZMHB) ; 5.X.1961, 1 ex ( ZMHB) ; Mähren , 2 exx ( NMW) ; 3 exx ( MTD); Mark Brandenburg, Schilow­Monchmuhle near Berlin, 25.IV.1984, Wrase, 1 ex ( ZMHB) ; Mittenwald , Bernhauer, 1 ex ( FMNH) ; München , 11 exx ( ZMHB) ; Neudorf , 3 exx ( NMW) ; Niederpoyritz , 3.VI.1907, 1 ex ( MTD) ; Ostrau, V. Zoufal, 2 exx ( NMW) ; Petersdorf See , 4 km S Furstenwalde, 2.I.1988, Uhlig, 1 ex ( ZMHB) ; Pferzheim environs, Enzgau, N. Baden , V–VI.1941, Netolitzky, 23 exx ( NMW) ; Potsdam , Glindow, 1 ex ( DEI) ; Potsdam, Hermannswerder Havel river , 23.IV.1978, Uhlig, 7 exx ( ZMHB) ; Scharpzow , 1863, 1 ex ( ZMUC) ; Schladen am Harz , 1 ex ( MHNG) ; Strausberg environs, 1 ex ( DEI) ; Stuttgart , Fischer, 1 ex ( CAS) ; Ulm , 15 exx ( MTD) ; Waltersdorf , Scheerpeltz, 1 ex ( NMW) ; Wernigerode, Bergwiese , 16.V.1980, D. Martin, 2 exx ( ZMHB) ; Wimpfen , 4 exx ( MNHN) ; Wismar, Hohen Wieschendorf, Tang , 12.VIII.1979, Uhlig, 2 exx ( ZMHB) ; Zeitz , 12–13.IX.1981, Uhlig, 1 ex ( ZMHB) ; “ Germany ” (localities not specified), 2 exx ( BMNH) ; 1 ex ( ZMUC); “ Saxonia ” (localities not specified), 1 ex ( MCSN) ; Markel , 1 ex ( MTD) ; “ Thuringia ” (localities not specified), 7 exx ( ZMHB) . Greece: Fthiotis, 30 km SW Lamia, Amfiklia, Kifisos river , 15.IV.2000, Assing, 1 ex ( VAPC) ; Grevena , 7.VI.1976, Sama, 3 exx ( AZPC) ; Kassandra, Sivirssa river , 23.III.1989, 2 exx ( VAPC) ; Pakas environs, 30.III.1986, 1 ex ( VAPC) ; Peloponnisos, 8 km NE Kalavrita, Vouraikos river , 700 m, 30.III.1997, Assing, 1 ex ( VAPC) . Hungary: Békés, Sarkad, Fekete­Körös , 90 m, 8.V.1982, Ádám, 2 exx ( HNHM) ; Békés, Sarkad, Remetei­erdö , 90 m, 24.V.1982, Ádám, 2 exx ( HNHM) ; Budafok., V. Stiller , 5.V.1917, 2 exx ( HNHM) ; Budapest, 23.V.1908, Rambousek, 1 ex ( NMPC) ; Bükk­hegys, Nagyvisnyó, Elzalak , 5–12.VI.1956, Kaszab & Székessy, 17 exx ( HNHM) ; Bükki National Park, Szarvasko , Eger, 250 m, 25.IV.1983, Ádám & Hámori, 1 ex ( HNHM) ; Bükki National Park, Cserépfalu, Hór­völgy , 250 m, 9.VII.1985, Ádám, 3 exx ( HNHM) ; Bükki National Park, Szarvasko , Eger, 250 m, 30.IV.1985, Ádám & Hámori, 10 exx ( HNHM) ; Bükki National Park, Miskolc, Hámori­tó , 300 m, 6.V.1981, Ádám & Migaly, 3 exx ( HNHM) ; Leányfalu , 21.4.1935, Csiki, 1 ex ( HNHM) ; Pilis­Hegyseg, Vörös­hegy near Piliscsaba , 5.VI.1980, Gollner, 2 exx ( ZMHB) ; Radvány , 8.5.1940, Csiki, 2 exx ( HNHM) ; Sopron, H., Moczarski, 1 ex ( NMW) ; Vas, Velem, Borha­forrás , 2.VI.1979, Ádám, 1 ex ( HNHM) . Italy: 50 km W Firenze, Padule di Fucecchio , 16.VI.1992, Assing, 3 exx ( VAPC) ; Abruzzes, Ovindoli val d'Arano , 1350 m, 3.VI.1952, G. Fagel, 6 exx ( IRSNB) ; Aguila , 30.IV.1904, A. Porta, 1 ex ( MSNM) ; Alta Vol Argentina, Realdo loc. Illin , 1300 m, 17.VIII.1974, Bologna, 1 ex ( AZPC) ; Anghebeni ( TN) , Leno river , 12.V.1992, Zanetti, 1 ex ( AZPC) ; Aspromonte, Calabria, Campi di Reggio , VI.1958, 1 ex ( MSNV) ; Aspromonte, Sant Eufemia , V.1957, 15 exx ( MSNV) ; Calabria, Santa Christina, Paganetti, 1 ex ( DEI) ; Carnia, Arta, Cedarchis , torr. Chiarso 400 m, 22.VI.1950, Fagel, 1 ex ( IRSNB) ; Cusercoli ( FO) , Fiume Ronco , 8.V.1976, Sama, 1 ex ( AZPC) ; Dolce ( VR) , Adige river , 25.VII.1976, Zanetti, 1 ex ( AZPC) ; Emilia, Coinacehio , 15.III.1901, Sangiorgi, 1 ex ( MSNM) ; Dora Baltea river, Ivrea , 9.VII.1958, Rosa, 1 ex ( MSNM) ; Feltre ( BL) , Rio Ligaut , 13.V.1973, 2 exx ( AZPC) ; Foce river , Torigno ( CB) , 28.VI.1976, W. Rossi, 1 ex ( AZPC) ; Gibilmanna , 900m, 19.5.1982, T. Palm, 1 ex ( MZLU) ; Giovi , VII.1905, 1 ex ( MSNM) ; Gubbio , 5.VIII.1891, 1 ex ( MSNM) ; La Varenna , 2 exx ( MNHN) ; Lazio­Riofreddo, Fosso Bagnatore , 1.VI.1943, M. Cerruti, 4 exx ( BMNH) ; Lombardia, Correzzana , 5.IV.1954, Brivio, 1 ex ( MSNM) ; 1.VI.1957, 8 exx ( MSNM) ; 9.VI.1957, 2 exx ( MSNM) ; 15.IX.1957, 1 ex ( MSNM) ; Lombardia, Brianza Case , VII.1959, Brivio, 3 exx ( MSNM) ; Lombardia, Monza­Parco , V.1957, Brivio, 3 exx ( MSNM) ; Lombardia, Monze M., IV.1961, Rose, 1 ex ( MSNM) ; Milano , 24.IV.1958, Rosa, 1 ex ( MSNM) ; Milano, Parco Monza , 13.IV.1957, Rosa, 1 ex ( MSNM) ; Montello , 6.IV.1955, 1 ex ( MSNM) ; Parco Lessinia, Valle Covoli di Velo , 22.VII.1991, Zanetti, 1 ex ( AZPC) ; Ponte Abbadia, affluente Fiora , 15.VII.1996, Zanetti, 3 exx ( AZPC) ; Revo Proves , 700–1400 m, 7.VIII.1996, Zanetti, 1 ex ( AZPC) ; Ronco all'Ad ( VR) , 25.IX.1971, Zanetti, 1 ex ( AZPC) ; Sicilia, Lake Pergusa, Enna , VI.1959, 5 exx ( MSNV) ; Stazzano Scrivia , 13.VI.1869, Ferrari, 3 exx ( MCSN) ; Trentino, Biotopo "La Rocchetta", 29.V.1996, 1 ex ( AZPC) ; Triest , Istrien, 1 ex ( NMW) . Poland: “Bellinchen” [Bielinek], 2 exx ( DEI) ; Beskid S decki, Majdan near Muszyna , 19.VII.2000, Melke, 1 ex ( AMPC) ; “ Beuthen ” [Bytom], Nowotny, 2 exx ( USMB) ; Bolechów­Stryj, Netolitzky , V.1917, 1 ex ( MHNG) ; “ Coeslin ”, Lullwitz, 1 ex ( USMB) ; Grochoszczyce , distr. Wadowice, Stobiecki, 1 ex ( ISEA) ; Jabłonna , 7.VII.1893, 5 exx ( USMB) ; Kazimierza Wielka , Klička, 4 exx ( NMPC) ; Kielcza ad Kędzierzyn Koźle, 2 exx ( USMB) ; Kraków , Rybiński, 1 ex ( ISEA) ; 3 exx ( NMW); Krzeszowice , distr. Chrzanów, Stobiecki, 1 ex ( ISEA) ; Łęczyca , 27.VII.1940, 2 exx ( MIZ) ; “ Legnitz ” [Legnica], Letzner, 6 exx ( DEI) ; Libertów , distr. Podgórze, Stobiecki, 1 ex ( ISEA) ; “ Nova Huta ” [Kraków­Nowa Huta], Lokay, 2 exx ( NMPC) ; Przemyśl , Kotula, 11 exx ( ISEA) ; Trella , 39 exx ( ISEA) ; “ Ratibor ” [Racibórz], H. Nowotny, 3 exx ( USMB) ; Rytro , distr. Nowy S cz, Stobiecki, 8 exx ( ISEA) ; “ Stettin ” [Szczecin], 1 ex ( NMW) ; “Zegrze” [ Warszawa­Zegrze ], 11.V.1935, Tenenbaum, 1 ex ( FMNH) . Russia: West Siberia , Barnaul, 1918–1920, H. Frieb, 6 exx ( NMW) ; Manina , 6.VI.1906, Klička, 5 exx ( NMPC) ; Moscow, Tschaschnikowo , 5.VII.1958, Hieke, 1 ex ( ZMHB) ; " Sarepta " [Volgograd], V. Bodemeyer, 1 ex ( FMNH) . Slovakia: Trenčin, 1 ex ( MTD) ; 1 ex ( ETH); Trenčin , 18.VII.1937, 1 ex ( ZMHB) . Spain: Cangas, 3 exx ( BMNH) ; Catalogne, Seo de Urgel, Arfa , V–VI.1962, Fagel, 6 exx ( IRSNB) ; Cueuca , 16.VI.1970, A. Comellini, 1 ex ( MHNG) ; G.C. Champion, 2 exx ( BMNH) ; La Granja , 1 ex ( BMNH) ; Madrid, Boadilla del Monte , NW Madrid, 6.VII.1996, Wrase, 3 exx ( MSPC) ; Ponferrada , Paganetti, 1 ex ( FMNH) ; Prov. Granada, Granada river near Rio Genil , 24.VII.1971, 1 ex ( MHNG) ; Seo de Urgel, Lerida , 15.VI.1969, Comellini, 7 exx ( MHNG) ; Soria, G.C. Champion, 2 exx ( BMNH) ; Valle de Saivoron, Espinana , 27.VII.1972, Puthz, 1 ex ( MHNG) . Sweden: Bosjoklaster , VII.1898, O. Moller, 3 exx ( MZLU) ; Flacharp, Trolleberg , 5.VI.1947, Sundholm, 1 ex ( MZLU) ; Hälsingborg , 15.V.1944, Palmqvist, 4 exx ( MZLU) ; 12.VI.1946, 1 ex ( MZLU) ; 12.V.1947, 1 ex ( MZLU) ; 15.V.1949, 1 ex ( MZLU) ; 18.VI.1950, Wiren, 5 exx ( MZLU) ; 18.VI.1956, E. Wiren, 4 exx ( MZLU) ; Hassleholm , 27.VII.1957, Israelson, 2 exx ( MZLU) ; Ivo , 2.VI.1947, A. Sundholm, 1 ex ( MZLU) ; Karlshamn , 28.V.1941, Sundholm, 1 ex ( MZLU) ; Karlskrona , 12.V.1942, A. Sundholm, 1 ex ( MZLU) ; Lomma , Palm, 2 exx ( MZLU) ; Lund , 11.V.1934, 4 exx ( MZLU) ; 13.VI.1935, Palm, 1 ex ( MZLU) ; 11.V.1939; 32 exx ( MZLU) ; Palm , 2 exx ( NHRS) ; Lund , 15 exx ( MZLU) ; Simrish , Palm, 1 ex ( MZLU) ; Vegeholm , 20.VI.1963, F. Olsson, 3 exx ( MZLU) ; Ven , V.1934, 1 ex ( MZLU) ; Vinslöv , 11.V.1959, Israelson, 1 ex ( MZLU) ; Vitemölla, Mollean , 21.VII.1947, Bo Tjeder, 1 ex ( MZLU) . Switzerland: Aaberg , Kanton Bern, V.1944, 2 exx ( ETH) ; Aarau , 1 ex ( ETH) ; Basel , 1 ex ( NHMB) ; 19.VI.1908, 1 ex ( NHMB) ; Bern, 3 exx ( NMW) ; “ Jhrich ”, 19.V.1909, 4 exx ( NHMB) ; II.1917, 2 exx ( NHMB) ; Morges , IV.1965, 1 ex ( NMW) ; Rhenian , 19.V.1910, 1 ex ( NHMB) ; Trient , V.1916, Notolitzky, 1 ex ( MHNG) ; Vaud, Anzeridaz , ca. 1800 m, 25– 31.VIII.1981, P. M. Hammond, 1 ex ( BMNH) ; Vaud, Solalex , ca. 1600 m, 25– 31.VIII.1981, P. M. Hammond, 1 ex ( BMNH) . Ukraine: Chlińska , distr. Ṡniatyń, Stobiecki, 2 exx ( ISEA) ; Drohobycz , Rybiński, 10 exx ( ISEA) ; Dźwinogród near Borszczów, 1 ex ( MIZ) ; Crimea, Yaila Region , Moczarski, 27 exx ( NMW) , Winkler , 2 exx ( DEI) ; Tarnopol, M. Rybiński, 1 ex ( ISEA) ; Zaleszczyki , 10.VII.1932, Tenenbaum, 1 ex ( FMNH) .

Redescription. Body. Length 3.2–3.5 mm, convex, parallel­sided, glossy; body colour pitchy brown to black; elytra brownish red, antennae testaceous to reddish brown, legs red to reddish brown, tarsi paler, yellow to reddish yellow, head and pronotum usually with brown reflection, three basal tergites often brownish red.

Head circular in outline, convex; eyes large, moderately protruding from lateral contours of head, length of each seen from above subequal to that of postocular region; surface of head without microsculpture; puncturation fine and very dense. Antennae long, clearly increased in width apically, antennomeres 2 and 3 subequal in length, antennomeres 4–7 longer than wide, antennomeres 8–10 quadrate, antennomere 11 nearly conical.

Pronotum quadrate, lateral sides gradually narrowed in straight line to obtuse hind angles; before base with small and shallow transverse impression; surface with fine isodiametric mesh microsculpture; puncturation fine, dense and asperate; pubescence at midline directed entirely anteriorly.

Elytra subquadrate, at suture as long as pronotum at midline; surface without microsculpture; puncturation fine, dense and asperate.

Abdomen parallel­sided, bases of tergites 3–5 each with deep transverse impression, impressions coarsely and densely punctuate, tergal puncturation uniformly fine and dense, tergite 8 with transversely stretched isodiametric mesh microsculpture.

Male. Aedeagus as in Figs. 19–20 .

Female. Spermatheca as in Fig. 21 .

Remarks. See under I. leucopus .

Distribution. Ischnopoda umbratica is widespread in the Palaearctic region. The distribution ranges from Spain in the west to the Ob River in the east.

Figures

FIGURES 19–21. Ischnopoda umbratica (Erichson): 19—aedeagus in lateral view, 20—aedeagus in ventral view, 21—spermatheca.