Carronella enne, Korshunova, Tatiana, Martynov, Alexander, Bakken, Torkild, Evertsen, Jussi, Fletcher, Karin, Mudianta, I Wayan, Saito, Hiroshi, Lundin, Kennet, Michael Schroedl, & Picton, Bernard, 2017

Korshunova, Tatiana, Martynov, Alexander, Bakken, Torkild, Evertsen, Jussi, Fletcher, Karin, Mudianta, I Wayan, Saito, Hiroshi, Lundin, Kennet, Michael Schroedl, & Picton, Bernard, 2017, Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda), ZooKeys 717, pp. 1-139 : 42-44

Carronella enne sp. n. Fig. 35

Type material.

Holotype, ZMMU Op-526, 5 mm long, North Atlantic W of Ireland, 48.7089'N, 10.5607'W, 11.06.2014, depth 1131 m, collected by Enrico Schwabe.

Type locality.

North Atlantic west of Ireland.

Etymology.

In honour of our friend, the malacologist Enrico (Enne) Schwabe (Bavarian State Collection of Zoology, Munich).

Diagnosis.

Mostly reduced notal edge, forming several clusters, background colour translucent white, digestive gland in cerata dull reddish, apical parts of cerata without white pigment, rachidian tooth with up to ten distinct denticles, adpressed to central cusp, lateral teeth smooth, penis conical.

Description.

External morphology (Fig. 35 A–E). Body relatively wide. Foot and tail narrow, anterior foot corners long. Rhinophores ca. 1.5 times longer than oral tentacles, slightly wrinkled. Dorsal cerata finger-shaped to fusiform, forming several clusters along dorsal edges. Apices of cerata gradually pointed, with elongate cnidosac. Distinct notal edge remains mostly below ceratal clusters and forms slight elevations. Digestive gland diverticulum fills significant volume of the cerata. Anal opening pleuroproctic on right side in between first and second large ceratal clusters. Reproductive openings lateral, below first anterior cluster of cerata.

Colour (Fig. 35A). Background colour translucent white. Digestive gland diverticula dull reddish. There are no opaque white lines on the body. Rhinophore background colour similar to body; a thin white line runs along dorsal side of rhinophores runs; apical white pigment absent. Apical parts of cerata with opaque cap of white pigment.

Jaws (Fig. 35G, H). Masticatory process more than one-third as long as jaw body. Edge of masticatory processes bears ca. 30 denticles that continue to form several reduced rows of denticles on body of masticatory processes.

Radula (Fig. 35I). Radula formula: 24 × 1.1.1. Rachidian tooth elongate-triangular with short narrow cusp of less than 1/3 of the tooth length (Fig. 35I). Rachidian tooth bears between eight and ten well-defined separated long lateral denticles strongly adpressed towards the cusp. Cusp is compressed by adjacent first lateral denticles. Lateral teeth (Fig. 35I) broadly triangular with obtuse and very attenuated posteriorly outer process, completely smooth.

Reproductive system (Fig. 35J). Diaulic. Hermaphroditic duct leads to convoluted ampulla of about two whorls. Vas deferens moderately long, no distinct prostate. Penial sheath small. Penis narrow conical. Oviduct connects through insemination duct into female gland complex. Vagina short and indistinct. Proximal receptaculum seminis oval. Distal receptaculum seminis present, small.

Ecology.

Deep sea species (deeper than 1000 m).

Distribution.

Northeast Atlantic, off Ireland.

Remarks.

According to the molecular phylogenetic analysis Carronella enne sp. n. forms a separate sister clade to Carronella pellucida (Fig. 1). Morphological analysis reveals differences in proportions of lateral teeth in C. enne sp. n. The lateral teeth of C. enne sp. n. are more elongated (Fig. 35I) than in C. pellucida (Fig. 36H).