Protohermes

Liu, Xingyue, Hayashi, Fumio & Yang, Ding, 2013, The Protohermes dichrous species group (Megaloptera: Corydalidae), with description of two new species from eastern Malaysia, Zootaxa 3620 (4), pp. 501-517 : 502

Genus Protohermes van der Weele

Protohermes van der Weele, 1907: 243. Type species: Hermes anticus Walker, 1853 , original designation.

Allohermes Lestage, 1927: 100 . Type species: Protohermes davidi van der Weele, 1909, original designation.

The Protohermes dichrous species group

Diagnosis. Body medium-sized (male wingspan ~ 70 mm). Head yellow, with a pair of black markings on vertex, postocular spine indistinct or absent. Posterior ocelli close to median ocellus. Wings hyaline, slightly brownish to grayish, with whitish or yellowish markings. Male ninth sternum posteriorly with a pair of digitiform or subtriangular lobes; ninth gonostylus straightly directed with base rather narrow; male ectoproct elongate and band-like, ~2.0–5.0 times as long as ninth tergum.

Remarks. Many species of the P. dichrous group in appearance resemble some species included in the P. sabahensis group by having similar vertex and wing marking patterns as well as some longitudinal veins with alternate color of black and yellow in the forewing. Although there is no phylogeny-based classification of the genus Protohermes , the straightly directed male ninth gonostylus with narrow base, which is a unique character in the P. dichrous group and the P. sabahensis group, may support the sister relationship between these two speciesgroups. The P. dichrous group is distinguished by the elongate and band-like male ectoproct versus a relatively short male ectoproct in the P. sabahensis group. However, whether the long male ectoproct is a reliable synapomorphy supporting the monophyly of the P. dichrous group is not confirmed.