Promalactis stenognatha Wang, 2021

Wang, Shuxia & Liu, Chen, 2021, Taxonomic study of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) X. The commotica species-group, with descriptions of twenty-two new species, Zootaxa 4980 (2), pp. 293-330 : 319-320

Promalactis stenognatha Wang , sp. nov.

( Figs 19 , 41 , 57 )

Type material. CHINA, Yunnan: Holotype ♂, Linjiapu (25.29°N, 98.70°E), Tengchong , 2144 m, 14.VIII.2014, leg. KJ Teng et al., slide No. JYY17593 . Paratypes (6♂, 6♀): 1♂, 6♀, same data as holotype, except 14‒16.VIII.2014, slide Nos. HXM14206 ♀, JYY17846 ♀, JYY17847 ♀, LC19775 ♂; 1♂, Nankang, Baoshan , 2009 m, 10.VIII.2014, leg. KJ Teng et al., slide No. JYY17861 ; 1♂, Nankang, Baoshan , 2009 m, 17.VII.2015, leg. KJ Teng & X Bai ; 3♂, Mt. Xiaohei, Longling , 1974 m, 28‒30.VII.2015, leg. KJ Teng & X Bai, slide Nos. JYY17583, JYY17582 .

Diagnosis. The new species can be distinguished by its narrow, slightly hooked gnathos, a costa with a large flagellate process, and a valva with two distal lobes curving in opposite directions.

Description. Adult ( Fig. 19 ). Forewing length 5.5‒6.5 mm.

Head: Vertex shiny white, frons greyish brown, occiput dim rust brown. Labial palpus with second palpomere orange yellow, paler on inner surface; third palpomere blackish brown except white at apex. Antenna with scape white; flagellum black, with white annulations on dorsal surface.

Thorax: Dorsum of mesothorax and tegula pale ocherous brown to rust brown. Forewing orange yellow, with blackish brown scales between fold and dorsum; markings white edged with blackish brown scales: costal spot inverted sub-triangular, extending obliquely outward to above posterior angle of cell; basal streak oblique inward to base of fold; antemedian streak from dorsal 1/3 to anterior margin of cell, arched outward medially; dorsal streak from basal 3/5 oblique outward to distal 1/4 of posterior margin of cell; tornal spot blackish brown, narrowed and diffused inward to end of dorsal streak; pretornal spot white, very small; apical spot rounded; two terminal spots smaller than apical spot, weakly separated by black scales; fringe orange yellow except greyish brown around tornus. Hindwing and fringe deep grey. Fore- and midlegs white on ventral surface; dorsal surface black, tibiae with two white stripes, with white scale tufts at apex, tarsi white at apices of basal two tarsomeres; hindleg yellow, tinged with blackish brown on dorsal side, tarsus white except second and third tarsomeres blackish brown at base.

Abdomen: Male genitalia ( Fig. 41 ). Uncus narrowed from base to narrowly rounded apex. Gnathos as long as uncus; mesial plate wide at base, distinctly narrowed and heavily sclerotized to pointed apex distally, slightly curved, with dense denticles; lateral arms wider and shorter than mesial plate. Tegumen furcate from posterior 1/3; lateral arms slightly narrowed to rounded apex. Valva relatively narrow, sub-parallel, with two distal lobes curving in opposite direction, forming a broad U-shape: dorsal lobe slender, extending dorsad, curved, rounded at apex, with a tooth at about middle; ventral lobe narrowed to apex, with an apical spine; costa with a long flagellate process arising from before dorsoapical process, far exceeding apex of dorsoapical process, with a longitudinal patch of long setae distally; sacculus ill-defined. Saccus slightly shorter than uncus, rounded anteriorly. Juxta broad, slightly widened to obtuse apex; basal lobe slender, longer than uncus. Aedeagus longer than valva, slightly curved, triangularly produced and with fine wrinkles distally, with fine denticles apically; cornutus absent.

Female genitalia ( Fig. 57 ). Apophyses posteriores 4 times length of apophyses anteriores. Lamella postvaginalis rectangular, longer than wide, notched at middle on posterior margin; lamella antevaginalis broad, deeply concave at middle on posterior margin, forming rounded posterolateral lobes with sparse setae, lateral margin with denticles along posterior half, concave at posterior 1/3. Antrum cup-shaped. Ductus bursae with basal 2/3 heavily sclerotized, slender, uniform; distal 1/3 broadened, membranous, with a sclerotized narrow coiled belt bearing 1‒2 short spines. Corpus bursae ovate, shorter than ductus bursae; signum an elongate ovate plate with two short denticles.

Distribution. China (Yunnan).

Etymology. The epithet of the new species is derived from the Greek steno- and gnathus, referring to the narrow hooked gnathos.

Figures

FIGURES 17‒22. Adults of Promalactis spp. 17, P. similibellatula, holotype, ♂; 18, P. squamacea, holotype, ♂; 19, P. stenognatha, holotype, ♂; 20, P. subgladiata, holotype, ♂; 21, P. tengchongensis, holotype, ♂; 22, P. triangulivalva, holotype, ♂. Scales = 2 mm.

FIGURES 40‒44. Male genitalia of Promalactis spp. 40, P. squamacea, holotype, slide No. DZH12334; 41, P. stenognatha, paratype, slide No. JYY17582; 42, P. subgladiata, holotype, slide No. LC19714; 43, P. tengchongensis, holotype, slide No. HXM14152; 44, P. triangulivalva, holotype, slide No. JYY17244. Scales = 0.2 mm.

FIGURES 56‒58. Female genitalia of Promalactis spp. 56, P. squamacea, slide No. DZH12245; 57, P. stenognatha, slide No. JYY17847; 58, P. subgladiata, slide No. LC19767. All paratypes. Scales = 0.2 mm.