Promalactis denticuligera Wang, 2021

Wang, Shuxia & Liu, Chen, 2021, Taxonomic study of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) X. The commotica species-group, with descriptions of twenty-two new species, Zootaxa 4980 (2), pp. 293-330 : 311-312

Promalactis denticuligera Wang , sp. nov.

( Figs 10 , 32 , 49 )

Type material. CHINA, Tibet: Holotype ♂ , Tongmai Town (30.10°N, 95.08°E), 2029 m, 12.VIII.2018, leg. MJ Qi, slide No. LC 19310 . Paratypes (23♂, 55♀): 6♂, 39♀, same data as holotype, except 12‒16.VIII.2019, slide No. LC 19311 ♀; 14♂, 5♀, same data as holotype, except 23‒26. VI.2009, leg. MJ Qi & JQ Deng, slide Nos. LC 19312 ♂, LC19313 ♀; 2♀, Yadong Village , Motuo County, 833 m, 16.VIII.2017, leg. XF Yang & MJ Qi, slide No. JYY17327 ; 1♂, 9♀, Pailong Town , Linzhi County, 2031 m, 17‒18.VIII.2018, leg. MJ Qi, slide Nos. LC 19326 ♂, LC19331 ♀; 1♂, Pailong Town , Linzhi County, 2010 m, 6.VII.2013, leg. JW Li, slide No. LC 19353 ; 1♂, Linzhi Town, Linzhi City , 2985 m, 19. VI.2019, leg. MJ Qi & JQ Deng, slide No. LC 19316 .

Diagnosis. The new species can be distinguished from other members of the group by two features of the male genitalia: the asymmetrically bilobed valvae, and the costa of the left valva produced into a free dorsodistal lobe bearing a cluster of spines along the costal margin.

Description. Adult ( Fig. 10 ). Forewing length 7.0‒9.0 mm.

Head: Frons greyish brown, vertex cream white, occiput rust brown. Labial palpus with second palpomere rust brown on outer surface, ochreous yellow on inner surface; third palpomere black on outer surface, rust brown on inner surface except white apically. Antenna with scape white dorsally, pale grey ventrally; flagellum black annulated with white on dorsal surface, black on ventral surface.

Thorax: Dorsum of mesothorax and tegula rust brown. Forewing ochreous brown on basal 2/3, ochreous yellow on distal 1/3; markings white edged with black scales: costal spot triangular, from distal 1/3 narrowed to above posterior angle of cell; basal streak slender, straight, oblique inward to base of fold; antemedian streak from dorsal 1/3 to anterior margin of cell, parallel with basal streak; dorsal streak from basal 2/3 extending obliquely outward to distal 1/4 of posterior margin of cell; tornal spot black, narrowly diffused inward to end of dorsal streak; distal patch black, extending from apex along termen to beyond tornal spot, gradually narrowed; apical spot sub-oval, small, set in black distal patch; fringe ochreous yellow except grey before tornus. Hindwing and fringe grey. Fore- and midlegs black, except fore femur white and midleg yellowish brown on ventral surface, tibiae with white spot at base and middle, with a tuft of white scales at apex, basal tarsomere white at base and apex, second tarsomere white apically; hindleg yellowish brown, tibia with long grey hairs, basal tarsomere white apically, second and third tarsomeres black except white distally, fourth and fifth tarsomeres white.

Abdomen: Male genitalia ( Fig. 32 ). Uncus wide parallel-sided from base to about basal 2/5, then slightly narrowed to rounded apex. Gnathos longer than uncus; mesial plate triangular, wide at base, narrowed to pointed apex, granulate distally; basal arms wide basally, narrowed to mesial plate. Tegumen with height more than three times length of uncus; lateral arms widened and arched on inner margin near apex. Valvae narrow at base, widened toward apex, asymmetrically bilobed distally; left valva: dorsal lobe slender, slightly narrowed to apex, ventral lobe wide at base, narrowed to basal 2/3, then slender to rounded apex, setose, shorter than dorsal lobe; costa very narrow, with clustered spines along costal margin in distal half; sacculus broadly banded, almost uniform in width, with a large semicircular concavity dorsally before free distal process; right valva: costa narrow, produced to a free digitate dorsal lobe distally, ventral lobe much larger and longer than dorsa lobe, elongate sub-triangular, with a flat apex; sacculus narrowly banded, wide at base, narrowed to basal 2/3, fused with valva and setose in distal 1/3. Saccus longer than uncus, elongate triangular, wide at base, narrowed to rounded apex. Juxta a large sclerotized plate, reaching posterior 1/4 of tegumen, narrow basally, widened distally, rounded apically; basal lobe clubbed, reaching middle of saccus. Aedeagus straight, almost as long as left valva, tubular, distal 1/3 spatulate, denticulate laterally and apically. Cornutus absent.

Female genitalia ( Fig. 49 ). Apophyses posteriores about 1/3 length of apophyses anteriores. Eighth sternum sub-rectangular, with a notch at middle on posterior margin.Antrum heavily sclerotized, slightly narrowed to ductus bursae. Ductus bursae membranous, slightly inflated, less than half length of antrum, with a coiled belt posteriorly. Corpus bursae slightly longer than ductus bursae, membranous, ovate; signum sub-ovate, with dense, large and small denticles.

Distribution. China (Tibet).

Etymology. The specific epithet is derived from the Latin denticuliger, referring to the distally and apically toothed aedeagus.

Figures

FIGURES 9‒16.Adults of Promalactis spp. 9, P. curvicosta, holotype, ♂; 10, P. denticuligera, holotype, ♂; 11, P. lianhuashana, holotype, ♂; 12, P. motuoensis, paratype, ♀; 13, P. pedata, holotype, ♂; 14, P. quadriprocessa, paratype, ♂; 15, P. ramiformis, paratype, ♂; 16, P. senticosa, holotype, ♂. Scales = 2 mm.

FIGURES 29‒34. Male genitalia of Promalactis spp. 29, P. binipapillata, holotype, slide No. JYY17611; 30, P. bispina, holotype, slide No. JYY17858; 31, P. curvicosta, holotype, slide No. LC19516; 32, P. denticuligera, holotype, slide No. LC19310; 33, P. lianhuashana, holotype, slide No. JYY17753; 34, P. motuoensis, holotype, slide No. JYY17672. Scales = 0.2 mm.

FIGURES 45‒50. Female genitalia of Promalactis spp. 45, P. arcicosta, slide No. LC19394; 46, P. binipapillata, slide No. JYY17673; 47, P. bispina, slide No. JYY17838; 48, P. curvicosta, slide No. LC19762; 49, P. denticuligera, slide No. LC19313, 49a, enlarged ductus bursae, 49b, enlarged singum, slide No. JYY17327; 50, P. motuoensis, slide No. JYY17597. All paratypes. Scales = 0.2 mm.