Eumeninae,

Ebrahimi, Ebrahim & Carpenter, James M., 2008, Catalog of the vespid wasps of Iran (Hymenoptera, Vespidae), Zootaxa 1785, pp. 1-42: 5-8

publication ID

http://doi.org/ 10.5281/zenodo.182436

persistent identifier

http://treatment.plazi.org/id/74608976-FFBC-FFA5-78E2-F8A0FA56F8A1

treatment provided by

Plazi

scientific name

Eumeninae
status

 

Subfamily Eumeninae  Leach

This is the most species rich subfamily in Iran, as indeed in the entire world. The following key allows separation of all the genera known from Iran. In the key, metasomal terga and sterna are denoted as “TI” for the first tergum, and so forth.

1. Metasoma petiolate: segment I in dorsal view with width half or less that of II, and at least twice as long as wide, usually longer ( Figs. 18View FIGURES 18 – 21, 23–25View FIGURES 22 – 25).................................................................................................... 2 - Metasoma not petiolate: segment I with width more than half that of II, much less than twice as long as wide ( Figs. 19–22View FIGURES 18 – 21View FIGURES 22 – 25, 26– 30View FIGURES 26 – 30)........................................................................................................................... 9 2. Forewing with recurrent veins received in separate cells ( Figs. 41View FIGURES 41 – 44, 45View FIGURES 45 – 48) ................... Raphiglossa  Saunders Forewing with recurrent veins both received in second submarginal cell ( Figs. 42–44View FIGURES 41 – 44, 46– 47View FIGURES 45 – 48) ................ 3 3. Midtibia with two spurs ( Fig. 48View FIGURES 45 – 48) .................................................................................. Discoelius  Latreille - Midtibia with one spur................................................................................................................................ 4 4. Propodeum posterolaterally toothed ( Fig. 67View FIGURES 67 – 68) .................................................. Cyrtolabulus  van der Vecht - Propodeum not toothed posterolaterally ( Fig. 68View FIGURES 67 – 68)....................................................................................... 5 5. TII with translucent apical lamella, clearly separated from disc by preapical thickening ( Figs. 9View FIGURES 6 – 10, 24View FIGURES 22 – 25) ........ ......................................................................................................................................... Eumenes Latreille  - TII with apical lamella not preceded by thickening ( Figs. 23, 25View FIGURES 22 – 25).............................................................. 6 6. TI much longer than mesosoma ( Fig. 23View FIGURES 22 – 25), parallel-sided for apical two thirds ( Fig. 18View FIGURES 18 – 21) and almost flattened dorsally ( Fig. 23View FIGURES 22 – 25); pronotum in dorsal view parallel-sided ...................... Ischnogasteroides  Magretti - TI not or little longer than mesosoma, apically swollen ( Fig. 25View FIGURES 22 – 25); pronotum in dorsal view convex anteriorly.............................................................................................................................................................. 7 7. Clypeus apically bluntly angular (female) or flatly convex (male; Fig. 35View FIGURES 35 – 40); tempora in dorsal view as long as eye; male with SVII broadly depressed.......................................................... Katamenes Meade-Waldo  - Clypeus apically emarginate ( Fig. 36View FIGURES 35 – 40); tempora in dorsal view shorter than eye; male with SVII not depressed..................................................................................................................................................... 8 8. Male TVII rounded and SVII with narrow median groove ( Fig. 69View FIGURES 69 – 71); TI usually smooth; male antenna with last article about as large as preceding articles................................................................ Delta  de Saussure - Male TVII hoodlike ( Fig. 71View FIGURES 69 – 71), SVII basally convex, distally forming lobate process ( Fig. 70View FIGURES 69 – 71); male antenna with last article reduced............................................................................. Alfieria Giordani Soika  9. Forewing with recurrent veins received in separate cells ( Figs. 41View FIGURES 41 – 44, 45View FIGURES 45 – 48) ..................... Psiliglossa  Saunders - Forewing with recurrent veins both received in second submarginal cell ( Figs. 42–44View FIGURES 41 – 44, 46– 47View FIGURES 45 – 48) .............. 10 10. Forewing with second submarginal cell petiolate anteriorly ( Fig. 47View FIGURES 45 – 48) ............................. Alastor  Lepeletier - Second submarginal cell not petiolate anteriorly ( Figs. 41–46View FIGURES 41 – 44View FIGURES 45 – 48) ............................................................... 11 11. Tegula evenly rounded posteriorly, not emarginate adjoining parategula and usually not reaching apex of latter ( Fig. 56View FIGURES 56 – 60); male antennae apically spiralled ...................................................................................... 12 - Tegula not evenly rounded posteriorly, emarginate adjoining parategula and usually reaching or surpassing apex of latter ( Figs. 57–63View FIGURES 56 – 60View FIGURES 61 – 63); male antennae apically hooked or simple ............................................. 19 12. Labial palpi 3 -segmented, in female broadly flattened or apically increasing in size, fringed with hairs, forming psammophore ( Figs. 31View FIGURES 31 – 34, 37, 39View FIGURES 35 – 40); male mandible with penultimate tooth often not excised....... 13 - Labial palpi 4 -segmented, female without psammophore ( Fig. 34View FIGURES 31 – 34); male mandible with penultimate tooth often deeply and broadly excised ............................................................................................................. 16 13. Forewing with prestigma longer than pterostigma ( Fig. 44View FIGURES 41 – 44); third segment of labial palpi not flattened; body pubescence short; female without cephalic foveae ........................................ Cephalochilus  Blüthgen - Forewing with prestigma shorter than pterostigma ( Figs. 41–42View FIGURES 41 – 44, 45– 47View FIGURES 45 – 48); third segment of labial palpi flattened or not; body pubescence long; female with cephalic foveae ........................................................... 14 14. Maxillary palpi five-segmented; labial palpi fringed with long setae apically, in female with both apical segments flattened ( Fig. 31View FIGURES 31 – 34); clypeal apex wide, about as wide as interantennal distance, truncate in female ( Fig. 31View FIGURES 31 – 34) and scarcely emarginate in male ........................................................... Pterocheilus  Klug - Maxillary palpi six-segmented; clypeal apex usually emarginate ............................................................ 15 15. Labial palpi not fringed with long hairs on apical half of third segment, in female both apical segments cylindrical, third pointed and needle-shaped ( Fig. 39View FIGURES 35 – 40); male without sternal brushes .................................. ................................................................................................................................. Hemipterochilus  Ferton - Labial palpi fringed with long hairs on apical half of third segment, in female third segment wide, flattened ( Fig. 37View FIGURES 35 – 40); male usually with dense brush of hairs on S III –V .............. Onychopterocheilus  Blüthgen 16. Prestigma longer than pterostigma ( Figs. 43–44View FIGURES 41 – 44); scutum with anterior smooth area .................................. ............................................................................................................................. Paravespa  Radoszkowski - Prestigma shorter than pterostigma ( Figs. 41–42View FIGURES 41 – 44, 45– 47View FIGURES 45 – 48); scutum without anterior smooth area … 17

17. Pronotum with sharply projecting anteroventral angle ( Fig. 55View FIGURES 51 – 55); parapsidal furrows distinct; mesepisternum with epicnemial carina; humeri often toothlike ........................................ Paragymnomerus  Blüthgen - Pronotum without sharply projecting anteroventral angle ( Fig. 51View FIGURES 51 – 55); parapsidal furrows absent .............. 18 18. Preoccipital carina present dorsally; female: cephalic foveae closely spaced, nearer occipital margin than posterior ocelli ( Fig. 38View FIGURES 35 – 40); clypeus deeply emarginate; mandible with penultimate tooth deeply excised; male: midlegs normal .............................................................................................. Gymnomerus Blüthgen  - Preoccipital carina absent dorsally; female: cephalic foveae well separated, located midway between posterior ocelli and occipital margin ( Fig. 40View FIGURES 35 – 40); clypeus truncate or weakly emarginate; mandible without excisions; male: midcoxae lobed, or midfemur ventrally with two excisions............... Odynerus Latreille  19. TI with transverse carina ( Figs. 19–21View FIGURES 18 – 21)..................................................................................................... 20 - TI without carina ( Figs. 22View FIGURES 22 – 25, 26–30View FIGURES 26 – 30) .......................................................................................................... 24 20 TI with broad longitudinal median furrow posterior to carina ( Fig. 20View FIGURES 18 – 21); mesepisternum with epicnemial carina; male antenna simple apically; female cephalic foveae well separated, located midway between posterior ocelli and occipital margin ( Fig. 40View FIGURES 35 – 40) ........................................................ Symmorphus Wesmael  - TI without broad groove ( Figs. 19, 21View FIGURES 18 – 21); mesepisternum without epicnemial carina; male antenna hooked apically; female cephalic foveae closely spaced, nearer occipital margin than posterior ocelli ( Fig. 38View FIGURES 35 – 40) 21 21. Submarginal carina not extended posteriorly as pointed process above valvula ( Fig. 50View FIGURES 49 – 50)............................ ................................................................................................................................. Ancistrocerus  Wesmael - Submarginal carina extended posteriorly as pointed process above valvula ( Fig. 49View FIGURES 49 – 50).............................. 22 22. Pretegular carina present ( Figs. 51, 55View FIGURES 51 – 55); epicnemial carina present or absent............................................... .............................................................................................................. Tachyancistrocerus  Giordani Soika - Pretegular carina absent ( Fig. 67View FIGURES 67 – 68); epicnemial carina present................................................................... 23 23. TI short, much wider than long; carina at about middle of tergum ( Fig. 19View FIGURES 18 – 21) Eustenancistrocerus Blüthgen  - TI about as long as wide or longer; carina at anterior narrow part of tergum, weak ( Fig. 21View FIGURES 18 – 21)...................... ........................................................................................................................... Jucancistrocerus Blüthgen  24. TII with translucent apical lamella, clearly differentiated from disc which is much thicker ( Figs. 26–29View FIGURES 26 – 30). .................................................................................................................................................................. 25 - TII without distinct lamella, sometimes bordered but apex not much thinner than disc ( Fig. 30View FIGURES 26 – 30)............ 26 25. TII without apical crenulae or foveae ( Figs. 28–29View FIGURES 26 – 30); propodeum with submarginal carina and vavula not projecting; scutum longer than wide ....................................................................... Microdynerus  Thomson - TII with apical row of crenulae or foveae ( Figs. 26–27View FIGURES 26 – 30); propodeum with submarginal carina and vavula differentiated; scutum variable, usually not or little longer than wide .................. Leptochilus  de Saussure 26. TI with transparent or translucent apical border ( Fig. 30View FIGURES 26 – 30); propodeum sometimes with superior carinae well developed, more or less lamelliform.................................................................................................. 27 - TI without border; propodeum without superior carinae........................................................................... 32 27. Propodeum without carinae or lamellae behind the metanotum (if superior carina present, it does not approach the scutellum)............................................................................................................................. 28 - Propodeum with carinae or lamellae behind the metanotum ( Figs. 74–75View FIGURES 72 – 75) .............................................. 29 28. Metanotum, seen from above, with semicircular ridge between the horizontal and vertical surfaces ( Fig. 73View FIGURES 72 – 75) ......................................................................................................................... Antodynerus  de Saussure - Metanotum, seen from above, with straight ridge between the horizontal and vertical surfaces, not arched ( Figs. 68View FIGURES 67 – 68, 72View FIGURES 72 – 75).......................................................................................................... Euodynerus Dalla Torre  [part: subgenus Euodynerus  ]

29. Parategula small, partly concealed by tegula; tegula narrow and apically bent inwards ( Fig. 59View FIGURES 56 – 60)................ .............................................................................................................................. Knemodynerus Blüthgen 

- Parategula large, not concealed by tegula; tegula shape different, apically not bent inwards ( Fig. 60View FIGURES 56 – 60).... 30

30. Pronotum without lamellate dorsal carina ( Fig. 84View FIGURES 81 – 87); epicnemial carina scarcely recognizable in general sculpture; metasoma dorsally covered with a short, recumbent bristles; male with midfemur flattened on 2 / 3 of the length and hindtibia distally swollen.................................................... Chlorodynerus Blüthgen 

- Pronotum with lamellate dorsal carina clearly developed ( Fig. 4View FIGURES 2 – 5); metasoma dorsally with little pubescence, only visible apically; male with midfemur not flattened and hindtibia not apically swollen......... 31

31. Metanotum not cristate; propodeum with superior carinae elevated near metanotum ( Fig. 74View FIGURES 72 – 75), more closely spaced than width of metanotum ( Fig. 75View FIGURES 72 – 75) ................................................ Syneuodynerus  Blüthgen

- Metanotum transverse, cristate along declivity ( Figs. 68View FIGURES 67 – 68, 72View FIGURES 72 – 75); propodeum with superior carinae not approaching metanotum so closely ........................................................................ Euodynerus  Dalla Torre [part: subgenus Pareuodynerus Blüthgen]

32. Pretegular carina absent ( Fig. 67View FIGURES 67 – 68)................................................................... Parodontodynerus Blüthgen 

- Pretegular carina present ( Figs. 51, 55View FIGURES 51 – 55) .................................................................................................... 33

33. Scutum posteriorly and scutellum impunctate.............................................................. Rhynchium Spinola 

- Scutum and scutellum punctate ................................................................................................................ 34

34. Metanotum bidentate ( Figs. 64–66View FIGURES 64 – 66)........................................................................................................... 35

- Metanotum not bidentate (in some Pseudepipona  slightly bilobed) ( Figs. 72–75View FIGURES 72 – 75)................................... 37

35. Metanotum laterally with short spines ( Fig. 66View FIGURES 64 – 66), flat in between.......................... Antepipona  de Saussure

- Metanotum laterally with strong upright teeth ( Fig. 65View FIGURES 64 – 66), depressed in between ( Fig. 64View FIGURES 64 – 66) ........................ 36

36. Scutellum bilobed, with deep transverse furrow anteriorly ( Fig. 64View FIGURES 64 – 66); propodeum not dentate; TII shallowly depressed .............................................................................................. Cyphodynerus  van der Vecht

- Scutellum evenly rounded; propodeum with several teeth laterally ( Fig. 65View FIGURES 64 – 66); TII not depressed ................ .................................................................................................................... Pseudodontodynerus Blüthgen 

37. Anterior face of pronotum with two small, close set, deeply impressed medial foveae ( Figs. 52, 54View FIGURES 51 – 55); tegula expanded semicircularly laterally ( Fig. 53View FIGURES 51 – 55); propodeum with submarginal carina produced into pointed lamella apically and valvula enlarged and free posteriorly from submarginal carina ............................... 38

- Anterior face of pronotum without two close set, deeply impressed foveae ( Figs. 51, 55View FIGURES 51 – 55); tegula variably shaped ( Figs. 61–63View FIGURES 61 – 63); submarginal carina and valvula usually not produced........................................... 39

38. Pronotum with foveae coalesced ( Fig. 52View FIGURES 51 – 55); TI subsessile, narrower than TII ............................................... ................................................................................................................... Paraleptomenes  Giordani Soika

- Pronotum with foveae separated ( Fig. 54View FIGURES 51 – 55); TI about as broad as TII ................. Stenodynerus  de Saussure

39. Propodeum with transverse ridge dorsally; tegula broadly rounded and expanded laterally, more than two thirds as wide as long ( Figs. 61–62View FIGURES 61 – 63) ......................................................................................................... 40

- Propodeum without transverse ridge dorsally; tegula narrow ( Fig. 63View FIGURES 61 – 63) .................... Allodynerus Blüthgen 

40. Tegula with coarse, sieve-like punctation ( Fig. 61View FIGURES 61 – 63); male mandible not notched ......................................... .......................................................................................................................... Brachypipona  Gusenleitner

- Tegula without coarse, sieve-like punctation ( Fig. 62View FIGURES 61 – 63); male mandible usually with a broad notch above the last tooth ........................................................................................................ Pseudepipona  de Saussure