Coprophanaeus (Coprophanaeus) morenoi Arnaud, 1982

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 69-71

Coprophanaeus (Coprophanaeus) morenoi Arnaud, 1982 Plate 16C

Coprophanaeus morenoi Arnaud, 1982b: 121 (original description. Type locality: Equateur [= Ecuador], km 36 route Sto Domingo-Quevedo).

Coprophanaeus morenoi : Vitolo 2000: 599 (characters in key); Medina et al. 2001: 140 (cited for Colombia); Arnaud 2002a: 42 (diagnosis); Vitolo 2004: 290 (redescription); Donoso et al. 2009: Appendix II. 16 (catalog of type MQCAZ); Edmonds and Zídek 2010: 57 (characters in key), 60 (redescription); Kohlmann and Solís 2012: 46 (cited for Ecuador); Krajcik 2012: 204 (complete list of species, cited as subgenus of Phanaeus Macleay, 1819);

Coprophanaeus (Coprophanaeus) morenoi : Carvajal et al. 2011: 320-321 (cited for Ecuador); Chamorro et al. 2018: 83 (figure 6C), 93 (cited for Ecuador).

Type specimens.

Coprophanaeus morenoi Arnaud, 1982. The holotype (♂) is deposited at the MNHN. Locality: Equateur [= Ecuador], Sto. Domingo km 36, examined. One Paratype is deposited in the MQCAZ, examined.

Holotype (♂): "EQUATEUR / STO DOMINGO K. 36 / JANV. 1982 / P&L ARNAUD coll [p and hw, black margin]", "Coprophanaeus / morenoi Mihi / HOLOTYPE ♂ / P. ARNAUD DET 1982 [p and hw, pink margin]", "HOLOTYPE [p, red label]".

Paratype (♀): "ECUADOR (PICH) / TINA LANDIA / I. 82 650 m / P & L ARNAUD leg [p]", "Coprophanaeus / morenoi Mihi / P. ARNAUD DET / PARATYPE ♀ [p and hw, pink margin]".

Distribution.

Colombia, Costa Rica, Ecuador, Nicaragua, and Panama.

Records examined.

CARCHI: Tobar Donoso, 300 m (8 specimens MECN). COTOPAXI: Guasaganda La Mana, 500 m (1 specimen CEMT; 3 specimens MQCAZ). ESMERALDAS: Colón del Onzole (17 specimens MECN); Charco Vicente (13 specimens MGO-UC; 28 specimens MECN); Gualpi (15 specimens MECN); Gualpi Pajonal (17 specimens MECN); Palma Real (1 specimen MGO-UC; 15 specimens MECN); Playa de Oro (1 specimen MUTPL; 10 speciemens MECN); Playa de Oro, Padre Santo (19 specimens MGO-UC; 31 specimens MECN); Playa de Oro, Estero Pote, 200 m (3 specimen CEMT; 16 specimens MECN); road Ibarra-San Lorenzo, El Placer, 670 m (1 specimen CEMT). PICHINCHA: Guayabilla Río Guayllabamba, Manduriacus, 520 m (2 specimens MUTPL); Llurimaguas Río Guayllabamba, Pedro Vicente Maldonado, 290 m (1 specimen MUTPL); Mangaloma, San Miguel de los Bancos, 820 m (1 specimen MUTPL); Tortugo Río Guayllabamba, Pedro Vicente Maldonado, 450 m (1 specimen MUTPL). SANTO DOMINGO DE LOS TSÁCHILAS: Tinalandia, 650 m (1 specimen MQCAZ).

Literature records.

ESMERALDAS: 11 km SE San Lorenzo, La Chiquita Sta, 5 m ( Edmonds and Zídek 2010: 62; Kohlmann and Solís 2012: 46); Yalare ( Edmonds and Zídek 2010: 62; Kohlmann and Solís 2012: 46); Punta Venado ( Edmonds and Zídek 2010: 62; Kohlmann and Solís 2012: 46); Playa de Oro ( Edmonds and Zídek 2010: 62; Kohlmann and Solís 2012: 46); La Concordia ( Edmonds and Zídek 2010: 62); Palma Real ( Edmonds and Zídek 2010: 62). GUAYAS: Los Ceibos ( Kohlmann and Solís 2012: 46); Borbón, 25 m ( Edmonds and Zídek 2010: 62); San Miguel ( Edmonds and Zídek 2010: 62). PICHINCHA: [= LOS RÍOS]: Station Río Palenque ( Arnaud 1982b: 122); 47 km S Sto. Domingo, Río Palenque ( Kohlmann and Solís 2012: 46); 113 km NW Quito on Puerto Quito road, 800 m ( Edmonds and Zídek 2010: 62). LOS RÍOS: Quevedo ( Edmonds and Zídek 2010: 62); Quevedo Pichilinge ( Kohlmann and Solís 2012: 46); Río Palenque Research Station, 200 m ( Edmonds and Zídek 2010: 62). SANTO DOMINGO DE LOS TSÁCHILAS: km 36 route Sto. Domingo-Quevedo ( Arnaud 1982b: 122); Tina Landia [= Tinalandia] (Env. Sto. Domingo) ( Arnaud 1982b: 122); i1 km E Tinalandia, 600 m ( Edmonds and Zídek 2010: 62) 16 km E Sto. Domingo, Tinalandia ( Kohlmann and Solís 2012: 46).

Temporal data.

Collected every month of the year.

Remarks.

Inhabits coastal lowland evergreen forests and coastal evergreen foothill forests from 5-850 m a.s.l. Species was collected with flight interception traps and pitfall traps baited with carrion and human feces.