Hadziella forneri, Quiñonero-Salgado & López-Soriano & Rolán, 2023

Quiñonero-Salgado, Sergio, López-Soriano, Joaquín & Rolán, Emilio, 2023, A new stygobiont species of the genus Hadziella Kuščer, 1932 (Gastropoda: Hydrobiidae) from Spain, Nemus 13, pp. 120-125 : 122-123

Hadziella forneri sp. n.

Type material: (Figs. 1-2). Holotype in MNHN (MNHN-IM-2000-38778) . Paratypes: 3 s in MNHN (MNHN-IM-2000-38779), 3 s in ZUPV-6840 5 s in SQS collection .

Type locality: Fuente de Morales , Calanda (Teruel province), Spain. [30TYL370357] . 381 m a.s.l.

This is a spring with abundant and permanent water flow, which rises in a small cavity and is channelized with a rubber pipe. It is hard to localize and to access, due to its abandoned state, and the amount of vegetation around. It is located in a slope close to the Guadalope River.

Etymology: The species is dedicated to Enric Forner i Valls, paleontologist specialized in sea urchins, and Editor of Nemus journal.

Description: Shell flat, planorbiform, compressed on both sides, lenticular and fragile. Very small in size, ranging from 0.23 to 0.32 mm in height and 0.70 to 0.83 in diameter. About 2 ½ whorls, from which 1.2 correspond to the protoconch. Marked sutures. Shell has a smooth appearance, but at high magnifications a microsculpture is evident in the protoconch, formed by many faint growth lines, slightly marked towards its last whorl, forming a rough surface at the end. In the umbilical part of the shell, a microsculpture formed by oblique lines irregularly arranged in spiral direction (Fig. 1F) is observed along the suture. Aperture oval-circular in shape with inclined axis, about 0.25 mm in diameter, with a thin border; in its inner part, which is detached from the previous whorl, a prominent area shows up which ends in a beak shape.

Dimensions: See Table 1 and Fig. 3 .

Habitat: Stygobitic. Shells were washed out from their habitat, likely after sudden rises of the subterranean water levels. Shells had a mainly fresh appearance, given its transparent color.

Distribution: Only known from the type locality.

Differential diagnosis: Shells of Hadziella forneri sp. n. can be distinguished from other species in the genus by these characters:

Hadziella anti Schütt, 1960 : Shell larger with more whorls (about 3 ½). A depression is observed both in upper and lower sides in the central part, while in the apical area there is a slight concavity. Apertural border is slightly tilted towards the outer side.

Hadziella rudnicae Bole, 1992 : Shell larger with more whorls, wider and more angled in the upper part, while slightly depressed in the lower part. It shows a slight concavity in the upper part.

Hadziella deminuta Bole, 1961 : Shell larger with more whorls (about 3 ½ - 4). Central part of both sides immersed, slightly wider the lower side, underside nearly flat. Upper apertural border slightly prolonged. Apical area showing a slight concavity in the upper part.

Hadziella sketi Bole, 1961 : Shell larger with more whorls. Slightly concave on both sides, and irregularly striated. Upper part of the peristome prolonged towards outside. Lower border of the shell angled. It shows a slight concavity in the upper part.

Hadziella thermalis Bole, 1992 : Shell larger with more whorls. Upper part slightly concave and regularly striated. Upper part of the peristome slightly elongated towards outside. In the apical area a slight concavity is present in the upper part.

Hadziella krkae Bole, 1992 : Shell larger with more whorls, slightly depressed on both sides. In the apical area a slight concavity is present in the upper part.

Hadziella ephippiostoma Kuščer, 1932 : Shell larger with more whorls (3-3 ½), slightly convex apical area.

Figures

FIGURE3.Measurements obtainedin the shells.SW:shelldiameter;SH: shellheight;PW:aperture width;PH:aperture height.