Vaquerosella perrillatae, Martínez-Melo & Alvarado-Ortega, 2020
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publication ID |
https://doi.org/ 10.26879/1040 |
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publication LSID |
lsid:zoobank.org:pub:5B28523A-C6D3-4AE1-AC2F-FF789BF99E4A |
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persistent identifier |
https://treatment.plazi.org/id/761AD07D-FF99-023E-7D45-F9EE0EA7F9C8 |
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treatment provided by |
Felipe |
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scientific name |
Vaquerosella perrillatae |
| status |
sp. nov. |
Vaquerosella perrillatae View in CoL sp. nov.
Figures 4-6 View FIGURE 4 View FIGURE 5 View FIGURE 6 ; Table 2.
zoobank.org/ 63CE8CF9-F1AC-4B54-B1C9-CA051527B929
Etymology. The specific name honors Dra. María del Carmen Perrillat, former curator of the Colección Nacional de Paleontología, Instituto de Geología, UNAM, in gratitude for her efforts to preserve the Mexican paleontological heritage.
Type material. Holotype IGM 11463 View Materials ( Figure 4 View FIGURE 4 ) . Paratypes IGM 11464 View Materials ( Figure 5.A View FIGURE 5 ) , IGM 11465 View Materials ( Figure 5.B View FIGURE 5 ) , IGM 11466 View Materials ( Figure 5.C View FIGURE 5 ) , IGM 11467 View Materials ( Figure 5.D View FIGURE 5 ) , IGM 11468 View Materials ( Figure 5.E View FIGURE 5 ) , IGM 11469 View Materials ( Figure 5.F View FIGURE 5 ) , and IGM 11470 View Materials ( Figure 5.G View FIGURE 5 ) . All specimens are from IGM-loc 248 except for IGM 11468 View Materials and IGM 11469 View Materials that were collected in IGM-loc 265. Additional material. IGM 11471 View Materials ( Figure 6.A, 6.C, 6.E View FIGURE 6 ) , IGM 11472 View Materials ( Figure 6.B View FIGURE 6 ) , IGM 11473 View Materials ( Figure 6.D View FIGURE 6 ) , IGM 11474 View Materials ( Figure 6.F View FIGURE 6 ) , IGM 11475 View Materials ( Figure 7.A View FIGURE 7 ) , IGM 11476 View Materials ( Figure 7.B View FIGURE 7 ) , IGM 11477 View Materials , IGM 11478 View Materials , IGM 11479 View Materials , IGM 11480 View Materials , IGM 11481 View Materials , IGM 11482 View Materials , and IGM 11483 View Materials . All specimens are from IGM-loc 248.
Type locality. IGM-loc 248, the Arroyo San Ignacio site, San Ignacio Formation, coordinates 27°09'22.00"N and 113°01'52.00"W, in the main drain of the San Ignacio River , Two paratypes are from the IGM-loc 265, Arroyo Nacho , located between the coordinates 27°09'22.00"N and 113°01'52.00"W. Both sites are into Mulegé Municipality, Baja California Sur, Mexico GoogleMaps .
Diagnosis. Test length and width nearly equal. Petal length three-fourths the distance from apical system to the ambitus. Ambitus with shallow indentations in each suture between columns of plates, resulting in a crenate margin. Anal notch small, slightly conspicuous, with soft projections on either side of posterior notch. Conspicuous notches at the ambulacra I and V, with soft projections on either side of lateral notch, acquiring a squared shape at the posterior margin.
Description. Test 46 to 77 mm in length ( Table 2). Aboral surface slightly domed, oral surface slightly concave. Highest point of test (7.2 to 15.7 mm) located at ambulacrum III, 34% of the length from the anterior edge. Outline slightly arrow-shaped from the aboral side, this due to the lateral notches in ambulacra I and V; with small notches in the other ambulacra. Test length almost equal to the width, with maximum width at interambulacra 1 and 4. Two very broad marginal indentations in ambulacra I and V, opening at an angle of about 120°. Very shallow anal notch, almost inconspicuous ( Figures 4-5 View FIGURE 4 View FIGURE 5 ).
Apical system monobasal, star-shaped, at 40- 50% L from the anterior edge of test; with numerous hydropores scattered over madreporic plate; four gonopores, one in each paired interambulacra and located at the suture between the madreporic plate and the first adapical plates of the interambulacral column ( Figure 6A View FIGURE 6 ).
Ambulacra petaloid, all approximately same length ( Figure 6E View FIGURE 6 ). Ambulacra I and V 75% of corresponding test radius; ambulacra II and IV 70% of corresponding test radius. Paired ambulacra mod-
MARTÍNEZ-MELO & ALVARADO-ORTEGA: NEW VAQUEROSELLA FROM MEXICO erately closed, maximum width at 45% of their length and close again to a 70% of their maximum width; with a few isolated pores outside petals. Anterior ambulacra petaloid III open, 75% of corresponding test radius, reaches its maximum width at the distal end ( Figure 4A View FIGURE 4 ). Petals slightly elevated; petals almost closed distally, with four to six trailing tube feet at distal end of each column of respiratory tube feet ( Figure 6E View FIGURE 6 ); not obviously lyrate, with outer pores tilted, with 6-7 tubercles in the poriferous zone ( Figure 6B View FIGURE 6 ).
The ambulacral pores conjugated, inner pore slightly oval-shaped, outer pore slit-like, comprising almost one third of pore pair, apparently subdivided by stereom septae ( Figure 6B View FIGURE 6 ). At ambitus, ambulacra greatly widened, length three times the width, curving strongly adapically to form test wall along each side of posterior notch. Ambulacral basicoronal plates all similar, narrow and straight with almost parallel radial sutures on each side ( Figure 6D View FIGURE 6 ).
Interambulacra narrow near the apical system, containing paired, zig-zag plates, interambulacral plates widened at ambitus, length 3x the width ( Figure 7 View FIGURE 7 ). Interambulacrum 5 not narrowing at ambitus but retaining same width as at end of petal.
On oral surface, three postbasicoronal plates in each half-interambulacrum in interambulacrum 5, three or four in the other interambulacra ( Figure 7 View FIGURE 7 ). All interambulacral basicoronals discontinuous, separated from first postbasicoronals by adjacent first ambulacral postbasicoronals, which are wider in interambulacrum 5 ( Figure 7B View FIGURE 7 ).
Peristome circular, relatively small (4-8% of L), anterior edge of peristome 45-50% L from the anterior edge of test, with distinct perradial process in each ambulacrum extending into peristome, beyond slight bulge containing sphaeridium ( Figure 6 View FIGURE 6 ). Anterior edge of peristome 54% TL from anterior edge of test. Food grooves well developed, restricted to oral surface, with no visible bifurcation near distal ends of ambulacral basicoronal plates. No significant depressions along perradial sutures on oral surface, no evidence of pressure drainage channels ( Figure 6B View FIGURE 6 ).
Periproct small (0.7-2% L), between third and fourth postbasicoronal interambulacral plates; usually on posterior margin but sometimes on oral side ( Figure 4B View FIGURE 4 ). Aboral tuberculation nearly homogeneous, with discernible enlargement of tubercles in oral interambulacral regions. Tube foot pores visible in food grooves.
Occurrence. IGM-loc 248 and IGM-loc 265, San Ignacio Formation, Mulegé, Baja California Sur, Mexico ( Figure 1 View FIGURE 1 ; Table 2).
Key for the Nominal Species of Genus Vaquerosella View in CoL
1. Test with a rounded ambitus, with no notches..2 - Test with postero-lateral notches, at the plates in ambulacra I and V......................3
2. Ambulacra wide open, specially the frontal ambulacrum........................... V. durhami View in CoL - Ambulacra opening until one-half the length, and then closing distally............. V. merriami View in CoL
3. Slight postero-lateral notches at the ambulacra I and V; ambitus rounded........... V. andersoni - Postero-lateral notches at the ambulacra I and V; ambitus with indentations at the perradial sutures. With projections in ambulacra 1, 4 and 5.... 4
4. Enlarged interambulacra 1, 4 and 5 ≥2x the length of ambulacrum I or V; this due to large projections at the interambulacra, giving the ambitus a stellate shape................................. 5
- Enlarged interambulacra 1, 4 and 5, <2x the length of ambulacrum I or V............... 6
5. Both frontal and paired petals open at the distal end................................ V. coreyi View in CoL - Frontal petal open but the paired petals closed at the distal end.............. V. vaquerosensis View in CoL
6. An almost triangular posterior margin due to deep notches at the posterior ambulacra and the projection of interambulacrum 5............ V. norrisi - A squared shape at the posterior margin, due to shallow posterior notches and two small projections besides the periproct...... V. perrillatae View in CoL
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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