Squatina

Walsh, Jonathan H., Ebert, David A. & Compagno, Leonard J. V., 2011, Squatina caillieti sp. nov., a new species of angel shark (Chondrichthyes: Squatiniformes: Squatinidae) from the Philippine Islands, Zootaxa 2759, pp. 49-59 : 58

Key to Western North Pacific Squatina Species (revised from Walsh & Ebert, 2007)

1a. Pelvic fin tips do not extend to origin of first dorsal fin........................................................ 2

1b. Pelvic fin tips extend to or surpass origin of first dorsal fin..................................................... 3

2a. A prominent row of thorn-like denticles extending from mid-back to caudal peduncle; no distinct ocelli on posterior lobes of the pectoral fins............................................................................... S. japonica

2b. No row of thorn-like denticles extending from the mid-back to the caudal peduncle, distinct paired ocelli on the posterior lobes of the pectoral fins....................................................................... S. tergocellatoides

3a. Dorsal-caudal space greater than inter-dorsal space; interorbital space greater than interspiracle space, pelvic inner margin greater than 9 % total length, pelvic base length greater than 13% total length......................................4

3b. Dorsal-caudal space less than interdorsal space; interorbital space less than interspiracle space, pelvic inner margin less than 9% total length, pelvic base length less than 13%.............................................. S. caillieti sp. nov.

4a. Upper lip arch semi-circular in shape (>1.5% TL in height); dorsals are lobed with a curvilinear anterior margin; pelvic girdle distance 1.4 times or less head length; caudal fin is lobed, especially dorsally, with a curvilinear postventral caudal margin............................................................................................... S. formosa

4b. Upper lip arch is not semi-circular in shape (<1.5% TL in height); dorsals are not lobed (angular) without a curvilinear anterior margin (straight); pelvic girdle distance greater than 1.4 times head length; caudal fin is not lobed (angular), especially dorsally, without a curvilinear postventral caudal margin (straight)......................................... S. nebulosa