Amblis Gistel, 1848

Bezděk, Jan, 2020, Review of the genus-level names proposed by Johannes Gistel in Chrysomelidae (Coleoptera), Acta Entomologica Musei Nationalis Pragae 60 (1), pp. 173-188 : 180-181

Amblis Gistel, 1848

Amblis Gistel, 1848a: 403 ; GISTEL (1848b): 123; GISTEL (1848c): [9]; GISTEL (1851): 123.

Type species. Chrysomela litura Fabricius, 1775, by monotypy.

Current status. Synonym of Spartophila Stephens, 1834 (subgenus of Gonioctena) (see MONRÓS & BECHYNÉ 1956, SEENO & WILCOX 1982, RILEY et al. 2003, KIPPENBERG 2010).

Comments. The name is also a junior homonym of Amblis Gistel, 1834 (a junior objective synonym of Chrysobothris Eschscholtz, 1829 , in Buprestidae ) (see BOUSQUET & BOU- CHARD 2017).

Chloemela Gistel, 1856, nomen oblitum Chloëmela Gistel, 1856a: 379; GISTEL (1856b): 175; GISTEL (1857a): 379.

Originally included names:

1. boica unavailable name, nomen nudum

2. aerea Me. = Chrysomela aerea Duftschmid, 1825 ; nomen dubium (see KIPPENBERG 2010), genus

Chrysolina Motschulsky, 1860 (Chrysome-

linae)

3. haematoptera (Lin.) = Chrysomela haemoptera Linnaeus, 1758; now Chrysolina (Colaphodes) haemoptera

(Linnaeus, 1758); genus Chrysolina Mo-

tschulsky, 1860 ( Chrysomelinae )

4. vulgatissima Schr. = Chrysomela vulgatissima Linnaeus, 1758; now Phratora vulgatissima (Linnaeus,

1758); genus Phratora Chevrolat, 1836

(Chrysomelinae)

5. sanguinolenta Lin. = Chrysomela sanguinolenta Linnaeus, 1758 ; now Chrysolina (Stichoptera) sanguinolenta

(Linnaeus, 1758); genus Chrysolina Mo-

tschulsky, 1860 ( Chrysomelinae )

6. limbata = Chrysomela limbata Fabricius, 1775; now Chrysolina (Zeugotaenia) limbata limbata

(Fabricius, 1775); genus Chrysolina Mot-

schulsky, 1860 ( Chrysomelinae )

7. marginata Lin. = Chrysomela marginata Linnaeus, 1758 ; now Chrysolina (Chalcoidea) marginata

marginata (Linnaeus, 1758) ; genus Chryso-

lina Motschulsky, 1860 ( Chrysomelinae )

8. analis Lin. = Chrysomela analis Linnaeus, 1767 ; now Chrysolina (Chalcoidea) analis (Linnaeus,

1767); genus Chrysolina Motschulsky, 1860

(Chrysomelinae)

9. lamina = Chrysomela lamina Fabricius, 1792; synonym of Chrysolina (Sulcicollis) oricalcia

(O. F. Müller, 1776); genus Chrysolina

Motschulsky, 1860 ( Chrysomelinae )

10. geminata Pay. = Chrysomela geminata Paykull, 1799; now Chrysolina (Hypericia) geminata (Paykull,

1799); genus Chrysolina Motschulsky, 1860

(Chrysomelinae)

11. viridula Lai. = Chrysomela viridula Laicharting, 1781; synonym of Chrysolina (Hypericia) hyperici

(Forster,1771); genus Chrysolina Motschul-

sky, 1860 ( Chrysomelinae )

12. varians = Chrysomela varians Schaller, 1783 ; now Chrysolina (Sphaeromela) varians (Schaller,

1783); genus Chrysolina Motschulsky, 1860

(Chrysomelinae)

13. fulgida = Chrysomela fulgida Fabricius,1801; synonym of Chrysolina (Euchrysolina) graminis

graminis (Linnaeus,1758); genus Chrysolina

Motschulsky, 1860 ( Chrysomelinae )

14. var. aurolimbata Bess. unavailable name, nomen nudum

15. graminis = Chrysomela graminis Linnaeus, 1758; now Chrysolina (Euchrysolina) graminis

graminis (Linnaeus,1758); genus Chrysolina

Motschulsky, 1860 ( Chrysomelinae )

16. fastuosa Lin. = Coccinella fastuosa Scopoli, 1763; now Chrysolina (Fasta) fastuosa fastuosa (Sco-

poli, 1763); genus Chrysolina Motschulsky,

1860 ( Chrysomelinae )

17. violacea = Chrysomela violacea Schaller, 1783; synonym of Chrysolina (Chrysomorpha) cerealis

megerlei (Fabricius, 1801); genus Chrysolina

Motschulsky, 1860 ( Chrysomelinae )

18. mixta Zie. = Chrysolina mixta Küster, 1844; synonym of Chrysolina (Chrysomorpha) cerealis me-

gerlei (Fabricius, 1801); genus Chrysolina

Motschulsky, 1860 ( Chrysomelinae )

19. cerealis Lin. = Chrysolina cerealis Linnaeus, 1767; now Chrysolina (Chrysomorpha) cerealis ce-

realis (Linnaeus, 1767); genus Chrysolina

Motschulsky, 1860 ( Chrysomelinae )

20. var. ericae Dahl = Chrysomela ericae Suffrian, 1851; synonym of Chrysolina (Chrysomorpha) cerealis

cerealis (Linnaeus, 1767); genus Chrysolina

Motschulsky, 1860 ( Chrysomelinae )

21. staphyleae Lin. = Chrysomela staphylaea Linnaeus, 1758 ; now Chrysolina (Chrysolina) staphy-

laea (Linnaeus, 1758); genus Chrysolina

Motschulsky, 1860 ( Chrysomelinae )

22. polita Lin. = Chrysomela polita Linnaeus, 1758; now Chrysolina (Erythrochrysa) polita

polita (Linnaeus, 1758); genus Chrysolina

Motschulsky, 1860 ( Chrysomelinae )

23. lurida = Chrysomela lurida Linnaeus, 1767 ; synonym of Chrysolina (Taeniosticha)

pseudolurida pseudolurida (Roubal, 1917);

genus Chrysolina Motschulsky, 1860 (Chry-

somelinae)

Type species. Chrysomela staphylaea Linnaeus, 1758 , by present designation.

Current status. Designated here as a nomen oblitum for Chrysolina Motschulsky, 1860 (nomen protectum).

Comments. The name is available based on Article 12.2.5 ( ICZN 1999). BOUCHARD et al. (2011) listed Chloemela Gistel, 1856 as a synonym of Chrysomela Linnaeus, 1758 . Because the type species was never designated and all included available species names belong to Chrysolina Motschulsky, 1860 and Phratora Chevrolat, 1836 , the synonymy with Chrysomela is incorrect.

GISTEL (1856a) listed altogether 23 names under the genus Chloemela. Two of these names, boica and var. aurolimbata , are nomina nuda, and thus are not available. The other 21 names are available and all but one of them are currently classified in the species-rich genus Chrysolina . The exception is C. vulgatissima classified in Phratora. I designate here Chrysomela staphylaea Linnaeus, 1758 as the type species of Chloemela.

The acceptance of Chloemela as a valid genus would destabilize the present system as widely used Chrysolina would become synonym of Chloemela. In order to preserve the stability, I apply Article 23.9 of the ICZN (1999) and designate Chloemela as a nomen oblitum and Chrysolina as a nomen protectum. To my knowledge, Chloemela was not treated as a valid name after 1899; thus, the conditions of Article 23.9.1.1 are met. According to Article 23.9.1.2 I supply the references of 25 papers, published by at least 10 authors in the immediately preceding 50 years, and encompassing a span of not less than 10 years, where Chrysolina Motschulsky, 1860 is used as a valid genus name: BIEŃKOWSKI (2007, 2008), BOURDONNÉ (1986), DACCORDI (1994), DACCORDI & RUFFO (1979, 1980), GEISER (2001), JOLIVET & HAWKESWOOD (1995), KIPPENBERG (2004, 2012, 2017), LOPATIN (1992, 2000, 2006), LOPATIN & KULENOVA (1986), MEDVEDEV (1999), MIKHAILOV (2000, 2001, 2002), RHEINHEIMER & HASSLER (2018), RILEY et al. (2003), WAR- CHAŁOWSKI (2010), WINKELMAN (2013), WINKELMAN & DEBREUIL (2008), YANG et al. (2015).