Edaphus permolestus, Puthz & D-, 2014

Puthz, Volker, 2014, Edaphus von Japan (Coleoptera: Staphylinidae) 118. Beitrag zur Kenntnis der Euaesthetinen, Revue suisse de Zoologie 121 (3), pp. 349-378 : 374-377

Edaphus permolestus sp. n. Abb. 46

HOLOTYPUS: MNSNT ohne Nummer ; ♂; JAPAN: Kyushu : Ohita pref., Mt. Kurodake, Kujyo, 16.IX.1985, S. Nomura .

PARATYPUS: NMSNT ohne Nummer; 1♂; Ohita pref., Yonouzu, 13. VI.1985, A. Miyata .

BESCHREIBUNG: Länge: 1,2-1,3 mm (Vorderkörperlänge: 0,6-0,65 mm). Apter, rötlichbraun, schimmernd, Stirn sehr fein punktiert, Elytren fein und dicht körnig skulptiert, restliche Oberseite bis auf die Insertionspunkte der anliegenden Beborstung unpunktiert.

PM des HT: HW:31; DE: 23; LE: 8; LT: 2: LG: 5; PW: 32; PL: 29; dlbc: 20,5; EW: 38,5; EL: 27; SL: 21.

Männchen: 4. Tergit mit sehr flachem Eindruck in der hinteren Mitte, 5. Tergit mit kaum erkennbarer Abflachung in der Mitte, beide fast borstenfrei und ohne dicht stehende Körnchen. 8. Sternit etwa wie in Abb. 23. 9. Sternit apikomedian spitz, mit zwei Wabenfeldern zu je etwa 16 ha. SpP und vs nicht gefunden. Aedoeagus (Abb. 46).

Dem E. permacer sp. n. sehr ähnlich, aber robuster, die Augen grösser, die Auszeichnung des 4. und Tergits beim ♂ anders.

BEMERKUNGEN: Von E perplexabilis sp. n. und E. perparvus sp. n., die beide auch am Mt. Kurodake erbeutet wurden, unterscheidet sich die neue Art so: vom

ersteren durch ihre Apterie, vom zweiten durch unpunktiertes Pronotum und die dicht und fein gekörnte Elytrenskulptur, von beiden durch die Sexualcharaktere des ♂. Zur Unterscheidung von den übrigen Arten vgl. die Bestimmungstabelle.

ETYMOLOGIE: Weil diese Art die Identifizierung der apteren Arten der aeneas -

Gruppe weiter erschwert, nenne ich sie „ permolestus “ (Lat. = sehr lästig“).

KEY TO THE EDAPHUS FROM JAPAN

[EXCLUSIVELY SOUTH JAPANESE (RYUKYU) ISLANDS]

1a Lateral furrows of frons distinct, becoming broader posteriorly near eyes (forming an impression, which may become as large as antennal segment 10 (Fig. 25) (exception: E. bishamon sp. n., where the lateral furrows are only angulated behind, slightly broadened).......................... 2

1b Lateral furrows of frons less distnct, very narrow, slightly curved (Figs 26, 27)................................................. 16

2a Mediobasal carina of pronotum absent or not extending to base proper..... 3

2b Mediobasal carina of pronotum extending to base proper................ 4

3a Lateral furrows of frons not broadened behind, antennal segment 10 distinctly broader than long. ♂: Abdominal tergites simple. ♂: E (Fig. 3). 1.3-1.5 mm (FB 0.7-0.8 mm). HON.................... bishamon sp. n.

3b Lateral furrows of frons becoming broader behind (about as in Fig. 25). E (Fig. 8). 1.2-1.4 mm (FB 0.7 mm).SHI.................. haniwa sp. n.

4a Mediobasal carina of pronotum short, not elongated anteriad, not extending to the middle of pronotum................................... 5

4b Mediobasal carina of pronotum longer, elongated anteriad, longer than half the length of pronotum (Figs 28, 30)............................ 9

5a Sutural length of elytra shorter than or at most as great as pronotal length. ♂: E (Fig. 6). 0.9-1.1 mm (FB 0.55-0.6 mm). SHI...... daimio sp. n.

5b Sutural length of elytra greater than pronotal length.................... 6

6a Elytra sparsely punctate. ♂: E (fig. 4, Puthz, 1975). 1.0-1.2 mm (FB 0.7 mm). HON, KYU............................. nipponensis Puthz

6b Elytra impunctate.............................................. 7

7a Anterolateral portions of frons with a distinct impression anteriorly. Pronotum with four large basal foveae. ♂: E (fig. 5, Puthz l. c.). 1.0-1.1 mm (FB 0.65-0.7 mm). HON, KYU.............. japonicus Sharp

7b Anterolateral portions of frons without a distinct impression anteriorly. Pronotum with six small basal foveae. ♂: E (figs 1-3, Puthz, l. c.). 0.91-1.1 mm (FB 0.6-0.7 mm). HON. Holarctic region .... lederi Eppelsheim

8a Brachypterous, sutural length of elytra less than pronotal length.......... 9

8b Macropterous, sutural length of elytra as long as or greater than pronotal length...................................................... 10

9a Mediobasal carina of pronotum very narrow, thread-like, elongated anteriad ♂: E (Fig.4). Elytra punctate. 1.0-1.2 mm (FB 0.6 mm). KYU................................................... sakura sp. n.

9b Mediobasal carina of pronotum, broadened anteriad, narrowly Y-shaped. ♂: E (Fig. 9). 1.0-1.2 mm (FB 0.6 mm). SHI, KYU.......... mikado sp. n.

10a Mediobasal carina very narrow, thread-like, elongated anteriad (Fig. 28).... 11

10b Mediobasal carina of pronotum less narrow, ± Y-shaped elongated anteriad (Fig. 30)............................................. 13

11a Smaller, FB 0.6 mm. Pronotum broader than long. ♂: E (fig. 6, Puthz, l. c.). 1.0-1.1 mm HON, SHI..................... carinicollis Bernhauer

11b Larger, FB 0.75-0.8 mm. Pronotum about as broad as long.............. 12

12a Anteromedian portion of frons flat; elytra very finely and densely punctate (setiferous punctures). ♂: E (Fig. 2). 1.2-1.4 mm. HON, KYU..................................................... samurai sp. n.

12b Anteromedian portion of frons callus-like elevated; elytra impunctate. ♂: E (Fig. 59). 1.2-1.5 mm (FB 0.7-0.8 mm).............. kanzeon sp. n.

13a Larger, FB 0.7-0.8 mm. E (Fig. 1). 1.2-1.4 mm (FB 0.7-0.8 mm). SHI, HON.............................................. shogun sp. n.

13b Smaller, FB ≤ 0.7 mm......................................... 14

14a Temples less than one quarter as long than eyes. ♂: E (Fig. 5). 1.1-1.5 mm (FB 0.6-0.7 mm). HOK, HON, SHI, KYU.. nakayamaensis Puthz

14b Temples longer than one quarter of eye length....................... 15

15a ♂: E (fig. 3, Puthz, 1980); Sternite 9 without fields of ha. 1.1-1.3 mm (FB 0.6-0.7 mm). HOK............................ napporoensis Puthz

15b ♂: E (Fig. 7), Sternite 9 with fields of 3-4 ha. 1.0 mm (FB 0.65 mm). HON............................................. bosatsu sp. n.

16a Lateral portions of frons with a minute impression anteriorly. Pronotum and elytra densely reticulate, slightly shiny. ♂: S8 (Fig. 16), SpP without any trumpet. 1.0 mm (FB 0.6 mm). HON................... tanuki sp. n.

16b Lateral portions of frons without an impression anteriorly. ♂ often with special sexual characters on tergites 4 and 5 (e. g. Figs 32, 35), SpP with a distal trumpet (e. g. Figs 39-41, 43-45, 47, 48).......... aeneas -group. 17

17a Macropterous................................................ 18

17b Micropterous or brachypterous, sutural length of elytra distinctly less than pronotal length, at most equal in length........................ 24

18a Pronotum and elytra with dense granular sculpture. No median carina on tergite 4. ♂: Unknown. 0.9-1.0 mm (FB 0.55. mm). KYU... persubtilis sp. n.

18b Pronotumand elytra indistinctly punctate........................... 19

19a ♂: Tergite 4 and 5 with median impressions......................... 20

19b ♂: Tergite 4 and 5 simple....................................... 21

20a ♂: E (Fig. 58). 1.2 mm (FB 0.6-0.65 mm). KYU, SHI.... perplexabilis sp. n.

20b ♂: E (Fig. 53). 1.0-1.1 mm (FB 0.6 mm). KYU......... percongruus sp. n.

21a Mediobasal carina of pronotum stronger, very distinct, mediobasal impressions of pronotum distinct, deeper........................... 22

21b Mediobasal carina of pronotum less strong, nearly indistinct, mediobasal impressions of pronotum shallower, less distinct..................... 23

22a ♂: E (Fig. 57). 1.3 mm (FB 0.7 mm). SHI.............. persimplex sp. n.

22b ♂: E (Fig. 56). 1.2-1.3 mm (FB 0.7 mm). KYU........ perpropinquus sp. n.

23a Anterolateral portions of frons impunctate. ♂: E (Fig. 55). 1.1-1.3 mm (FB 0.7-0.75 mm) SHI.............................. peramicus sp. n.

23b Anterolateral portions of frons finely punctate. ♂: Unknown. 1.1-1.2 mm (FB 0.7 mm). HON................... perincommodus sp. n.

24a Pronotum punctate............................................ 25

24b Pronotum (besides of setiferous punctures) impunctate................. 29

25a Elytra impunctate. ♂: Tergite 4 with a distinct impression posteromedially, tergite 5 with a shallow median impression (Fig. 35), E (Figs 49, 50). 1.2-1.3 mm (FB 0.55-0.6 mm). KYU............. perparvus sp. n.

25b Elytra punctate or granulose..................................... 26

26a Elytra punctate............................................... 27

26b Elytra granulose. ♂: Tergite 4 and 5 with very shallow impressions medially. Posteromedian callus of frons very small. 1.1-1.3 mm (FB 0.55 mm). HON................................. perangustus sp. n.

27a Punctation of elytra denser, interstices smaller than punctures. ♂: Ter- gites 4-6 with very shallow impressions medially, E (about as in Fig. 51). 1.1-1.3 mm (FB 0.65 mm). SHI....................... pergracilis sp. n.

27b Punctation of elytra less dense, interstices larger than punctures. ♂: Tergites 4 and 5 with distinct impressions medially................... 28

28a E (Fig. 54). 1.3-1.4 mm (FB 0.65-0.7 mm). HON......... perillustris sp. n.

28b E (Fig. 51). 1.4 mm (FB 0.75 mm). HON................. perversus Puthz

29a Larger, FB ≥0.7 mm. ♂: Tergites 4 and 5 with distinct impressions medially, E (Fig. 52). 1.4-1.5 mm (FB 0.75 mm). HON...... pergratus sp. n .

29b Smaller, FB 0.5-0.6 mm........................................ 30

30a Eyes smaller, elytra very shallowly, densely sculptured. ♂: Tergites 4 and 5 with an area of dense granulae, E (about as in Fig. 51). 1.2-1.4 mm (FB 0.5-0.6 mm). HON.............................. permacer sp. n.

30b Eyes larger, elytra densely and finely granulose. ♂: Tergite 4 with a very shallow apicomedial impression, tergite 4 and 5 with an area of less dense pubescence medially. E (Fig. 46). 1,2-1,3 mm (FB 0.6-0.65 mm). KYU.......................................... permolestus sp. n.