Orasema evansi Burks, Heraty & Dominguez, 2018

Burks, Roger A., Heraty, John M., Dominguez, Chrysalyn & Mottern, Jason L., 2018, Complex diversity in a mainly tropical group of ant parasitoids: Revision of the Orasema stramineipes species group (Hymenoptera: Chalcidoidea: Eucharitidae), Zootaxa 4401 (1), pp. 1-107 : 33-47

Orasema evansi Burks, Heraty & Dominguez n. sp.

urn:lsid:zoobank.org:act:2CBC91DD-A3EF-44FD-B0C1-C1C7605DE6D2 ( Figs 2 , 61–76 )

Orasema sp. C1 near costaricensis ; Heraty 1990, 1994b.

Diagnosis. Mesoscutal midlobe and mesoscutellar disc mostly finely reticulate. Mesoscutellar disc with fine sculpture posteromedially. Axillula with parallel carinae and no other sculpture.

This species is most similar to O. costaricensis , O. minutissima and O. rapo . It differs from O. costaricensis and O. minutissima in lacking reticulate sculpture on the axillula ( Figs 65 , 73 ). It differs from O. rapo in having an areolate-reticulate or reticulate, instead of wholly areolate, mesoscutellum ( Figs 64 , 72 ). These differences are slight, but are supported by molecular data.

Description. Female. Length 3.4–4.9 mm.

Color: Head and mesosoma green to purple, with varying luster. Pedicel yellow; anellus brown; flagellum brown. Coxae brown; fore femora yellow or with brownish coloration; mid and hind femora yellow. Fore wing slightly infuscate; venation pale brown. Petiole purple; gaster brown with green to purple luster.

Head: Head width:head height 1.1–1.2; face reticulate; longitudinal groove between eye and torulus absent; eyes bare, inter-ocular distance:eye height 1.4–1.5; malar space:eye height 0.6–0.9; supraclypeal area longer than broad, smooth to weakly sculptured; clypeus smooth. Flagellum length:head height 1.4–1.5; F2 2.5–3.0× as long as broad, 1.1–1.2× as long as F3; following funiculars subequal in width, progressively shorter.

Mesosoma: Mesosoma length:height 1.4. Mesoscutal midlobe reticulate to weakly areolate-reticulate or posteriorly smooth, with minute setae; lateral lobe areolate-reticulate to nearly smooth. Axilla areolate-reticulate; mesoscutellar disc areolate-reticulate; frenal line regularly foveate; frenum areolate-reticulate; axillular groove anteriorly absent, broad and foveate posteriorly; axillula with longitudinal carinae. Propodeum with irregular median carina; propodeal disc areolate to areolate-reticulate; callus reticulate, with 3–5 setae. Mesepisternum reticulate to areolate-reticulate laterally, smooth ventrally, broadly rounded anterior to mid coxa. Upper mesepimeron smooth; lower mesepimeron reticulate; transepimeral sulcus distinct. Metepisternum laterally smooth to reticulate. Hind coxa 2.3–2.4× as long as broad; hind femur 5.4–5.5× as long as broad. Fore wing 2.4– 2.6× as long as broad, 2.7–3.0× as long as mesothorax; stigma with 4 or 5 sensilla in an irregular row. Hind wing costal cell with a broad bare area.

Metasoma: Petiole 2.4–2.6× as long as broad, 0.9–1.0× as long as hind coxa, areolate, anterior carina weak. Antecostal sulcus crossed by carinae; acrosternite posteriorly angulate.

Male. Length 2.2–2.3 mm. Head width:head height 1.2; scape yellow; flagellum length:head height 1.7–2.0, F2 1.8–2.4× as long as broad. Fore femur yellow or with a brownish area, mid and hind femora, and all tibiae yellow. Petiole 5.5–6.2× as long as broad, petiole length:hind coxa length 1.8–1.9.

Discussion. The series of this species found ovipositing into bananas was previously reported as Orasema costaricensis by Kerrich (1963). These specimens, which were morphologically agreeable with one another in having a yellow fore femur and sculptured mesoscutal sidelobes, were designated as holotype and paratypes. Many other specimens with a smooth sidelobe were also present in this series, and were indistinguishable from specimens associated with Alchornea costaricensis Pax & K.Hoffm. (Euphorbiaceae) instead of bananas. We have not been able to obtain sequence data from specimens found ovipositing on either of these plants. Morphologically comparable specimens are tentatively included in this species but were not designated as paratypes. We also have not been able to consistentlyl distinguish specimens that we morphologically attribute to O. evansi from Central America and the Caribbean from specimens in the United States ( Fig. 2 , inset) despite their very different behaviors (oviposition into bananas of soft leaves of A. costaricensis versus hard leaves of Quercus (Fagaceae) or Myrica (Myricaceae) .

Biology. The holotype and paratypes were reported as ovipositing into bananas ( Fig. 3 ) by Evans (1968). Orasema evansi (as O. costaricensis ) was considered enough of a pest in the 1970s that insecticidal treatments were used in an attempt to control the host ants ( Ostmark & Evers 1976; Wojcik 1990). A series of planidia were collected in Coto, Costa Rica from leaves of mango and citrus that were likely part of the surveys of bananas at that time (see Other Material Examined). While these planidia may not be definitively assignable to O. evansi , they are morphologically similar and may attest to a much broader host plant range. Some tentatively assigned (not holotype or paratype) Costa Rican specimens were collected on leaves of A. costaricensis , assigned to O. costaricensis based on morphology.

The Texas and Florida populations were reared from Pheidole dentata ( Van Pelt 1950; Heraty 1990, 1994b). Adults were observed ovipositing onto the underside of leaves of small plants of water oak, Quercus nigra L. ( Fagaceae ) in Texas (JMH). In Texas (Stubblefield Lake), the host colony of P. dentata was found in the soil directly below the host plant in a mixed pine oak forest. Van Pelt reared them from a colony collected in a decayed log of Pinus palustris Mill. (Pinaceae) in a stand of longleaf pine flatwoods. The rate of parasitism of P. dentata was reported by Heraty (1994b), referring to this species as Orasema sp. [C1]. Parasitism rates are difficult to measure given the differnt larval stages and potential inclusion of phthisergates, but were as high as 53.5–54.2% of the brood ( Heraty 1994b).

Immature stages (Texas population, Fig. 4 ). Egg white, smooth and stalked. ( Fig. 4A ). First instar ( Figs 4B, D, E ). Unfed planidium length 0.26 mm; after the planidium burrows into the host larva, the first instar swells and the tergites are reduced to bands on the sac-like larva ( Fig. 4D ); shape is retained after the larval exuvium is shed ( Fig. 4E ) and the larva migrates to a ventral position between the legs of the host pupa. In all other respects, the planidium is the same as that of Orasema delicatula (Walker) (Burks et al. 2015: Fig. 17 ): antenna absent; labial plates absent; tergopleural line absent; tergites I and II (TI, TII) separate; TI–III with dorsal seta and with wrinklelike longitudinal sculpture; TII and TVI with very short lateral seta; TI, TIII, TV, TVII with ventral seta; TIX with separated leaflike ventral plate having a uniformly curved posterior margin; caudal pad present; caudal cerci present and relatively long, about as long as apical 4 tergites. Second instar ( Fig. 4F ). Smooth dorsally and weakly sculptured ventrally; single thoracic spiracle. Third instar and Prepupa ( Figs 4G, H ). Swollen with lateral tubercles dorsal and ventral to the seven lateral spiracles. Prepupa with six ventral transverse bands of weak sculpture; lateral tubercles and dorsal body segments papillate. Pupa ( Fig. 4I ). Head and mesosoma lacking tubercles; three dorsal tubercles over petiolar region; mesosoma with raised lateral bands and posteriorly with paired dorsal tubercles.

Distribution. Costa Rica, Dominican Republic, Honduras, Nicaragua, Panama, St. Vincent, Trinidad, USA (Virginia to Texas).

Material examined. Holotype: HONDURAS: La Lima , 15°08'00"N, 85°52'00"W, 24.v.1963, C. Evers, oviposition on bananas [1 ♀, UCRCENT00248573 ], deposited in USNM . Paratypes: COSTA RICA: Alajuela: Chiles de Aguas Zarcas, Café , 300m, 10°23'56"N, 84°20'20"W, x.1989, R. Cespedes [1 ♀, UCRC: UCRCENT00237534 ] . R.N.V.S. Caño Negro, 20m, 10°53'32"N, 84°49'57"W, 19–27.ii.1993, K. Flores [2, INBIO: INBIOCRI01193262, INBIOCRI01193164]. Alajuela: Playuelas, R.N.V.S. Caño Negro , 20m, 10°53'32"N, 84°47'12"W, 10–31.x.1992, K. Flores [2 ♀, INBIO: INBIOCRI00943984 , INBIOCRI00944047] . Guanacaste: Nandayure , 0–5m, 9°48'06"N, 85°17'34"W, 23.vi.2004, M. Moraga, Manglar Jabilla, red Noyes [2 ♀, INBIO: INB03983106 , INB03983107 ] . Heredia: La Selva 3km S Pto. Viejo , 10°26'00"N, 84°01'00"W, 6.iv.1985, H.A. Hespenheide [1 ♀, UCRC: UCRCENT00243573 ] . La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 7.v.1990, H.A. Hespenheide [1 ♀, UCRC: UCRCENT00446973 ] . La Selva Biol. Sta. 3km S Pto. Viejo , 10°26'00"N, 84°01'00"W [1 ♀, UCRC: UCRCENT00316334 ] . Puntarenas: Coto , 9°33'00"N, 84°09'00"W, 5.iii.1956, R.V. Roig, on bananas [8 ♀, USNM: UCRCENT00248622–25 , UCRCENT00248635–38 ] . Coto, Golfito, 9°39'04"N, 83°56'53"W, 11.i.1956, Jose Varella, on bananas fingers [5 ♀, UCRC: UCRCENT00479163; USNM: UCRCENT00248626–28, UCRCENT00248633]. N.E. 65, C.E. Palma [1 ♀, USNM: UCRCENT00248634 ] . Rancho Quemado, Peninsula de Osa , 200m, 8°40'39"N, 83°33'53"W, vii.1991, F. Quesada [1 ♀, INBIO: INBIOCRI01408645 ] . HONDURAS: La Lima , 15°08'00"N, 85°52'00"W, 24.v.1963, C. Evers, oviposition on bananas [72 ♀, USNM: UCRCENT00248174 , UCRCENT00248176 , UCRCENT00248182–83 , UCRCENT00248189 , UCRCENT00248197 , UCRCENT00248199 , UCRCENT00248201 , UCRCENT00248204– 205 , UCRCENT00248533–72 , UCRCENT00248574–84 , UCRCENT00248586–94 ,

UCRCENT00248616–17]. La Lima , 15°08'00"N, 85°52'00"W, 16.vii.1959, R. Bullock, Ovip. on bananas [1 ♀, USNM: UCRCENT00248618 ] . 460m, 15°08'00"N, 85°52'00"W, 13.iv.1963, E. Melendez [4 ♀, UCRC: UCRCENT00237491–93, UCRCENT00237640]. La Lima , 15°08'00"N, 85°52'00"W, 24.v.1963, C. Evers, oviposition on bananas [25 ♀, BMNH: UCRCENT00309887–910 , UCRCENT00309912 ] . La Ceiba: 15°45'38"N, 86°46'51"W, F.J. Dyer [2 ♀, USNM: UCRCENT00248585, UCRCENT00248615]. NICARAGUA: Granada: Baker [1 ♀, USNM: UCRCENT00248027 ] . PANAMA: Gamboa , 9°07'00"N, 79°42'00"W, 10.v.1972, R & E Froeschner [1 ♀, USNM: UCRCENT00248643 ] . Montebrio , iii.1924, D.T. Fullaway [1 ♀, BPBM: UCRCENT00429465 ] .

Additional material examined: COSTA RICA: Alajuela: Chiles de Aguas Zarcas, 300m, 10°23'56"N, 84°20'20"W, i.1990, R. Cespedes, cafe [5 ♀, UCRC: UCRCENT00251472, UCRCENT00251482, UCRCENT00252339, UCRCENT00252347, UCRCENT00252349]. Chiles de Aguas Zarcas, 300m, 10°23'56"N, 84°20'20"W, iii.1990, R. Cespedes, cafe [15 ♀, UCRC: UCRCENT00414276–85, UCRCENT00414287–88, UCRCENT00414290–91, UCRCENT00414293]. Chiles de Aguas Zarcas, Café, 300m, 10°23'56"N, 84°20'20"W, x.1989, R. Cespedes [5 ♀, UCRC: UCRCENT00251468–71, UCRCENT00251480]. Chiles de Aguas Zarcas, Café, 300m, 10°23'56"N, 84°20'20"W, xi.1989, R. Cespedes [4 ♀, CNC: UCRCENT00425962–64; MUCR: UCRCENT00479299]. Chiles de Aguas Zarcas, Café, 300m, 10°23'56"N, 84°20'20"W, xii.1989, R. Cespedes [8 ♀, MUCR: UCRCENT00479305, UCRCENT00479310, UCRCENT00479312; UCRC: UCRCENT00251473–74, UCRCENT00252342, UCRCENT00252350–51]. Finca La Selva NE Dos Rios, 400m, 10°24'48"N, 84°29'35"W, 27.iii.1988, Hanson [1 ♀, UCRC: UCRCENT00252338]. Heredia: Chilamate, 75m, 10°26'60"N, 84°05'55"W, 25.ii.1989, P. Hanson [1 ♀, UCRC: UCRCENT00252341]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 1.iv.1987, H.A. Hespenheide [3 ♀, UCRC: UCRCENT00243554–56]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 1.iv.1987, H.A. Hespenheide, on leaves of Alchornea Pax Hoffm. [3 ♀, LACM: UCRCENT00242172, UCRCENT00242174, UCRCENT00242191]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 11.iv.1988, H.A. Hespenheide, on leaves of Alchornea Pax Hoffm. [1 ♀, LACM: UCRCENT00242190]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 13.iv.1988, H.A. Hespenheide [3 ♀, UCRC: UCRCENT00446627, UCRCENT00446788, UCRCENT00446984]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 2.iv.1987, H.A. Hespenheide [5 ♀, UCRC: UCRCENT00243560, UCRCENT00243572, UCRCENT00243567–68, UCRCENT00243561]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 2.v.1990, H.A. Hespenheide [1 ♀, UCRC: UCRCENT00446972]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 23.iii.1987, H.A. Hespenheide, on leaves of Alchornea Pax Hoffm. [3 ♀, LACM: UCRCENT00242176–78]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 24.iii.1987, H.A. Hespenheide, on leaves of Alchornea costaricensis Pax Hoffm. [11 ♀, LACM: UCRCENT00242175, UCRCENT00242166–70, UCRCENT00242181– 82, UCRCENT00242184–86]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 26.iii.1987, H.A. Hespenheide, on leaves of Alchornea Pax Hoffm. [2 ♀, LACM: UCRCENT00242197–98]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 27.iii.1987, H.A. Hespenheide, on leaves of Alchornea Pax Hoffm. [1 ♀, LACM: UCRCENT00242183]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 28.iii.1987, H.A. Hespenheide [8 ♀, UCRC: UCRCENT00243546, UCRCENT00243550, UCRCENT00243551, UCRCENT00243553, UCRCENT00243559, UCRCENT00243562, UCRCENT00243571, UCRCENT00279853]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 28.iii.1987, H.A. Hespenheide, on leaves of Alchornea Pax Hoffm. [4 ♀, LACM: UCRCENT00242171, UCRCENT00242180, UCRCENT00242193, UCRCENT00242194]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 29.iii.1987, H.A. Hespenheide, Hamelia [4 ♀, UCRC: UCRCENT00243557–58, UCRCENT00243570, UCRCENT00243576]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 29.iii.1987, H.A. Hespenheide, on leaves of Alchornea Pax Hoffm. [2 ♀, LACM: UCRCENT00242195–96]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 30.iii.1987, H.A. Hespenheide [2 ♀, UCRC: UCRCENT00243563, UCRCENT00243569]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 31.iii.1987, H.A. Hespenheide [6 ♀, UCRC: UCRCENT00243548, UCRCENT00243564–66, UCRCENT00279852, UCRCENT00243547]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 31.iii.1987, H.A. Hespenheide, on leaves of Alchornea Pax Hoffm. [2 ♀, LACM: UCRCENT00242187–88]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 31.v.1991, H.A. Hespenheide [1 ♀, UCRC: UCRCENT00446628]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 4.v.1990, H.A. Hespenheide [1 ♀, UCRC: UCRCENT00446983]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 6.iv.1988, H.A. Hespenheide, on leaves of Alchornea Pax Hoffm. [1 ♀, LACM: UCRCENT00242192]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 7.v.1990, H.A. Hespenheide [3 ♀, UCRC: UCRCENT00446971, UCRCENT00446981–82]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 8.may.1990, H.A. Hespenheide [1 ♀, UCRC: UCRCENT00446979]. La Selva 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 9.iv.1987, H.A. Hespenheide, on leaves of Alchornea Pax Hoffm. [1 ♀, LACM: UCRCENT00242173]. La Selva Biol.Sta. 3 km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 9.iv.1989, H.A. Hespenheide [2 ♀, UCRC: UCRCENT00243574, UCRCENT00243578]. La Selva Biol. Sta. 3 km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 17.vii.1992, H.A. Hespenheide [1 ♀, UCRC: UCRCENT00446976]. La Selva Biol. Sta. 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W [8 ♀, UCRC: UCRCENT00243581, UCRCENT00316319–22, UCRCENT00316332–33, UCRCENT00316335]. La Selva Biol. Sta. 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 1.v.1990, H.A. Hespenheide, on leaves of Alchornea costaricensis [2 ♀, UCRC: UCRCENT00400742, UCRCENT00446980]. La Selva Biol. Sta. 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 27.iv.1990, H.A. Hespenheide, on leaves of Alchornea costaricensis [1 ♀, UCRC: UCRCENT00446977]. La Selva Biol. Sta. 3km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 30.iv.1990, H.A. Hespenheide, on leaves of Alchornea costaricensis [4 ♀, UCRC: UCRCENT00279851, UCRCENT00446974–75, UCRCENT00446978]. La Selva Biol. Sta., 3 km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 1.v.1990, H.A. Hespenheide [1 ♀, UCRC: UCRCENT00479166]. La Selva Biol. Sta., 3 km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 13.iv.1988, H.A. Hespenheide [1 ♀, UCRC: UCRCENT00479165]. La Selva Biol. Sta., 3 km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 17.viii.1996, H.A. Hespenheide [1 ♀, UCRC: UCRCENT00412935]. La Selva Biol. Sta., 3 km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 19.vi.1991, H.A. Hespenheide [1 ♀, UCRC: UCRCENT00400741]. La Selva Biol. Sta., 3 km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 5.iv.1988, H.A. Hespenheide, on leaves of Alchornea costaricensis Pax & Hoffm. [2 ♀, UCRC: UCRCENT00243575, UCRCENT00243577]. La Selva Biol. Sta., 3 km S Pto. Viejo, 10°26'00"N, 84°01'00"W, 5.viii.1996, H.A. Hespenheide [1 ♀, UCRC: UCRCENT00412936]. La Selva Field Station near Puerto Viejo, 10°25'53"N, 84°00'13"W, 21–28.iii.1988 [1 ♀, UCRC: UCRCENT00416651]. Puerto Viejo La Selva, 100m, 10°25'00"N, 84°01'00"W, 21.i–3.ii.1991, J. Noyes [1 ♀, UCRC: UCRCENT00414274]. Puerto Viejo, La Selva, 100m, i.1991, J. Noyes [1 ♀, MUCR: UCRCENT00479309]. Limón:, 10°06'23"N, 83°30'25"W, xi.1959, N.L.H. Krauss [1 ♀, USNM: UCRCENT00248653]. Puntarenas: Coto, Golfito, 9°39'04"N, 83°56'53"W, 11.i.1956, Jose Varella, on bananas fingers [1 ♀, BMNH: UCRCENT00412718]. DOMINICAN REPUBLIC: Santa Domingo, 18°28'44"N, 69°53'27"W, 16.i.1989, L. Masner, urban riparian, screen sweep [2 ♀, CNC: UCRCENT00247565–66]. PANAMA: Bocas del Toro: Changuinola, 9°29'17"N, 82°33'37"W, xii.1967, C.S. Stephens, on bananas, pan trap [2 ♀, USNM: UCRCENT00248646–47]. Coiba: Estacion Biologica , 24.i.1994, J.L. Nieves, sweep [1 ♀, MNCN: UCRCENT00456009]. 2 km S Torti, Serrania de Maje, 8°53'00"N, 78°24'00"W, 18.i.2001, L. Masner, ss [4 ♀, UCRC: UCRCENT00091459–61, UCRCENT00235925 (D1603)]. ST. VINCENT: Leeward side, 555m, 13°15'11"N, 61°10'05"W, H.H. Smith [1 ♀, BMNH: UCRCENT00309911]. Mayreau Island, 50m, 12°38'33"N, 61°23'35"W, 13–17.viii.2009, S. Peck, thorn scrub forest, MT [1 ♀, CNC: UCRCENT00425822 (D4648)]. TRINIDAD: 30.xi.1928, Myers [1 ♀, BMNH: UCRCENT00309913]. St. George: Curepe, 10°38'48"N, 61°24'56"W, 11.ii.1979, malaise trap [1 ♀, CNC: UCRCENT00247574]. Curepe, 10°38'48"N, 61°24'56"W, 7.vii.1978, malaise trap [1 ♀, CNC: UCRCENT00247575]. Dept. Agriculture grounds, Port of Spain, 10°39'34"N, 61°28'44"W, 24.x.1918, Harold Morrison [1 ♀, USNM: UCRCENT00248595]. Curepe, CIBC lab grounds, 10°38'48"N, 61°24'56"W, 14–29.iii.1974, E.D Bennett [1 ♀, CNC: UCRCENT00247571]. Curepe, CIBC lab grounds, 10°38'48"N, 61°24'56"W, 6.xii.1974 – 30.i.1975, F.D Bennett & S. Singh [1 ♀, CNC: UCRCENT00247570]. Curepe Sta. Margarita, Circular Rd., 10°39'14"N, 61°24'08"W, 8–15.xii.1975, F. Bennett, pan trap [1 ♀, FSCA: UCRCENT00241373]. Curepe, 10°38'48"N, 61°24'56"W, 13.iii.1979, Malaise trap [1 ♀, CNC: UCRCENT00425709]. Curepe, CIBC grounds, 10°38'20"N, 61°23'54"W, 15–29.iv.1974, M.N. Beg, moericke trap [1 ♀, CNC: UCRCENT00425707]. Curepe, CIBC lab grounds, 10°38'48"N, 61°24'56"W, 14– 24.iii.1974, M.N. Beg, Moericke trap [1 ♀, CNC: UCRCENT00425706]. Curepe, Santa-Margarita Circular Road, 10°39'30"N, 61°24'12"W, 1.ix.1974, F.D. Bennett [3 ♀, CNC: UCRCENT00425703–05]. Curepe, Santa-Margarita Circular Road, 10°39'30"N, 61°24'12"W, 15–25.v.1978, F.D. Bennett [1 ♀, CNC: UCRCENT00425701]. Mt. St. Benedict Trail, 330m, 10°40'01"N, 61°24'02"W, 16–17.vii.2013, Heraty & Baker, ypt [1 ♀, UCRC: UCRCENT00412117 (D3799)]. St. Augustine, vii.1976, J.S. Noyes, Moericke Trap [1 ♀, BMNH: UCRCENT00309459]. St. Augustine, 10°38'19"N, 61°23'46"W, ix.1976, F.D. Bennett [2 ♀, BMNH: UCRCENT00309462–63]. St. Augustine, 10°38'19"N, 61°23'46"W, 15.vii–15.viii.1976, J.S. Noyes, malaise trap [4 ♀, UCRCENT00309455–58]. Curepe, 10°38'48"N, 61°24'56"W, 23.iii.1979, malaise trap [1 ♀, CNC: UCRCENT00425708]. Curepe, CIBC lab grounds, 10°38'48"N, 61°24'56"W, 13.vii–31.viii.1974, M. N. Beg [1 ♀, CNC: UCRCENT00247573]. Curepe, Sta. Margarita, Circular Rd., 10°39'14"N, 61°24'08"W, 28.i–9.ii.1974, E.D Bennett [1 ♀, CNC: UCRCENT00247572]. Curepe, CIBC lab. grounds, 10°38'20"N, 61°23'54"W, 15.iv.1974, F.D. Bennett [1 ♀, CNC: UCRCENT00425702]. St. Augustine, 10°38'19"N, 61°23'46"W, ix.1976, F.D. Bennett [2 ♀, BMNH: UCRCENT00300431–32]. St. George Co., Curepe, 10°38'38"N, 61°24'40"W, ix.1976, F.D. Bennett [1 ♀, BMNH: UCRCENT00300434]. St. Augustine, 10°38'19"N, 61°23'46"W, 18.vi.1976, J.S. Noyes, wasteground [3 ♀, BMNH: UCRCENT00300441–43]. St. Augustine, 10°38'19"N, 61°23'46"W, 24.vi.1976, J.S Noyes [1 ♀, BMNH: UCRCENT00300440]. St. Augustine, 10°38'19"N, 61°23'46"W, 15.vii–15.viii.1976, J.S. Noyes, malaise trap [3 ♀, BMNH: UCRCENT00300424–25, UCRCENT00300428]. St. Augustine, 10°38'19"N, 61°23'46"W, 19.vi.1976, J.S. Noyes, wasteground [1 ♀, BMNH: UCRCENT00300444]. St. Augustine, 10°38'19"N, 61°23'46"W, 21.vi.1976, J.S. Noyes, wasteground [2 ♀, BMNH: UCRCENT00300439, UCRCENT00300445]. USA: Florida: Lacoochee Fla., 28°27'57"N, 82°10'19"W, 18.viii.1930, L.D. Tuthill [1 ♀, UCRC: UCRCENT00252352]. Cedar Key, 29°08'19"N, 83°02'06"W, 29.viii.1976, Z. Bouček [1 ♀, BMNH: UCRCENT00309922]. Orange Co., UCF, Orlando, MacKay Tract, 28°35'27"N, 81°13'01"W, 8.ii.1999, P. Russell, S. Fullerton, Sawgrass Marsh/ Red Maple, MT [1 ♀, UCFC: UCFC00014326]. Orange Co., Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 1–14.xi.1994, J.C. Longhurst, S.M. Fullerton, S Pine / Oak Scrub, MT [1 ♀, UCFC: UCFC00165849]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 12.vii.1995, S.M Fullerton, Summer Student, S Pine / Oak Scrub, MT [2 ♀, UCFC: UCFC00165860, UCFC00165875]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 17.viii.1995, S.M. Fullerton, Summer Student, S Pine / Oak Scrub, MT [3 ♀, UCFC: UCFC00165841, UCFC00165847, UCFC00165854]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 20.xi.1994, J.C. Longhurst, S.M. Fullerton, S Pine / Oak Scrub, MT [1 ♀, UCFC: UCFC00165837]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 24.ix.1994, J.C. Longhurst, S.M. Fullerton, S Pine / Oak Scrub, MT [3 ♀, UCFC: UCFC00165861–62, UCRCENT00446684]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 27.viii.1994, J.C. Longhurst, S.M. Fullerton, S Pine / Oak Scrub, MT [1 ♀, UCFC: UCFC00165871]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 5.vii.1995, S.M. Fullerton, Summer Student, S Pine / Oak Scrub, MT [2 ♀, UCFC: UCFC00165863, UCFC00165876]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 5.viii.1995, S.M. Fullerton, Summer Student, S Pine / Oak Scrub, MT [2 ♀, UCFC: UCFC00165839, UCFC00165848]. Orange Co., Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 3.i.1995, J.C. Longhurst, S.M. Fullerton, S Pine /Scrub Oak, MT [1 ♀, UCFC: UCFC00032719]. Orange Co., Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 11.v.1995, S.M. Fullerton, Spring Students, S Pine / Oak Scrub, MT [1 ♀, UCFC: UCFC00165844]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 12.vii.1995, S.M. Fullerton, Summer Student, S Pine / Oak Scrub, MT [1 ♀, UCFC: UCFC00165874]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 19.vii.1995, S.M. Fullerton, Summer Student, S Pine / Oak Scrub, MT [1 ♀, UCFC: UCFC00165842]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 28.vi.1995, S.M. Fullerton, Summer Student, S Pine / Oak Scrub, MT [1 ♀, UCRC: UCRCENT00446683]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 7.vi.1995, S.M. Fullerton, Summer Student, S Pine / Oak Scrub, MT [2 ♀, UCFC: UCFC00165843, UCFC00165870]. Cedar Key, 29°08'19"N, 83°02'06"W, 29.viii.1976, Z. Bouček [2 ♀, UCRC: UCRCENT00251465–66]. Eureka nr. Ocala, 29°22'20"N, 81°54'16"W, 3.ix.1976, Z. Bouček [2 ♀, BMNH: UCRCENT00239257–58]. Fakkahatchee, 26°01'12"N, 81°24'00"W, xii.1975, Marcio Naves [1 ♀, TAMU: UCRCENT00184061]. Gainesville A.E.I, 29°39'06"N, 82°19'29"W, 14–26.xi.1986, D. Wahl [1 ♀, EMUS: UCRCENT00247581]. Gainesville A.E.I, 29°39'06"N, 82°19'29"W, 5.xii.1986, D. Wahl [1 ♀, CNC: UCRCENT00247582]. Gainesville Am. Ent. Inst., 29°39'30"N, 82°22'46"W, 20.xi.1986, G. Gibson, sweep [1 ♀, CNC: UCRCENT00247583]. Gainesville, 29°39'06"N, 82°19'29"W, 28.viii.1976, Z. Bouček [1 ♀, UCRC: UCRCENT00251467]. Gainesville, 29°39'06"N, 82°19'29"W, 6–14.x.1986, D. Wahl [1 ♀, CNC: UCRCENT00247580]. Jacksonville, 30°19'56"N, 81°39'20"W [1 ♀, USNM: UCRCENT00248620]. Jacksonville, St. Johns Bl., 30°15'53"N, 81°23'51"W, 30.viii.1976, Z. Bouček [2 ♀, BMNH: UCRCENT00239259, UCRC: UCRCENT00251463]. Osceola, Walt Disney World Conservation Area, 28°18'13"N, 81°30'26"W, 5–19.ix.1994, Z. Prusak, MT [1 ♀, UCFC: UCFC00165877]. Sanford, 28°47'59"N, 81°16'42"W, M.C. Van Duzee [1 ♀, UCRC: UCRCENT00237630]. Alachua Co., Gainesville A.E.I., 33m, 29°35'53.6"N, 82°21'54.8"W, 13–22.ix.2003, D.B. Wahl [2 ♀, CNC: UCRCENT00320783 (D3938), UCRCENT00320784 (D3957)]. Gainesville EMUS, 29°39'08"N, 82°22'25"W, 13–23.x.1997, L. Masner [5 ♀, CNC: UCRCENT00091316–19, CNC: UCRCENT00235959]. Gainesville, 29°39'06"N, 82°19'29"W, 13.vi.1996, L. Davis [1 ♀, FSCA: UCRCENT00411928]. Gainesville, 29°39'06"N, 82°19'29"W, P. Hibbs [2 ♀, UCRC: UCRCENT00439144–45]. Gainesville, 29°46'38"N, 82°27'17"W, 12.x.1946, A. Van Pelt [1 ♀, USNM: UCRCENT00248621]. Gainesville; EMUS, 29°35'56"N, 82°21'55"W, 22–25.xi.1996, L. Masner, pan trap [2 ♀, CNC: UCRCENT00425829–30]. Baker Co., Glen St. Mary, 30°18'00"N, 82°00'55"W, 20.ix.2007, E. Zoll, S. Fullerton, rural yard/mixed woods, malaise [1 ♀, UCFC: UCFC00394715]. Glen St. Mary, 30°18'00"N, 82°00'55"W, 28.vi.2007, E. Zoll, S. Fullerton, rural yard/mixed woods, malaise [1 ♀, UCFC: UCFC00392628]. Brevard Co., Titusville, SR 405, Enchanted Forest Sanct. White Trail, 5m, 28°32'00"N, 80°48'09"W, 11–25.vii.2001, P.J. Russell, Z. Prusak and S.M. Fullerton, Xeric Oak Hammock, Malaise Trap [1 ♀, UCFC: UCFC00118065]. Colombia & Baker Co., Osceola N.F., 28°47'35"N, 81°03'33"W, 26.ix–8.x.1976, J.R. Wiley, malaise trap [1 ♀, FSCA: UCRCENT00241374]. Holly Hill, 29°14'37"N, 81°02'15"W, 1.vi.1978, G.J. Umphrey [3 ♀, UCRC: UCRCENT00237563, UCRCENT00237567–68]. Seminole Co., Lower Wekiva River St. Preserve, Burn Zone LW-5, 28°49'28"N, 81°24'26"W, 17.v.1999, P.J. Russell and S.M. Fullerton, Longleaf Pine—Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00111591], 21.i.2001, P.J. Russell and S.M. Fullerton, Longleaf Pine-Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00107788]. Osceola N.F., RT. 90, 28°47'35"N, 81°03'33"W, 8–20.x.1976, J.R. Wiley, malaise trap [1 ♀, FSCA: UCRCENT00241378]. Dade Co., 25°48'30"N, 80°17'09"W, 14.ii.1978, L.A. Stange [1 ♀, FSCA: UCRCENT00241383]. Duval Co., St. Johns Bluff, 30°23'06"N, 81°30'02"W, 30.viii.1976, E.E. Grissell [1 ♀, FSCA: UCRCENT00241379]. Highlands Co., Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 1– 29.ii.1988, D.B. Wahl, malaise trap [1 ♀, UCRC: UCRCENT00251450]. Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 1.vi–16.vii.1987, D.B. Wahl, malaise trap [1 ♀, CNC: UCRCENT00247577]. Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 12.xi.1998, M. Deyrup, below Pinus clausa with Cinara aphids, yellow pan trap [1 ♀, ABS: UCRCENT00305270]. Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 18– 19.x.1998, M. Deyrup, west side of LK. Annie, area of chironomid and spider abundance, yellow bowl trap [1 ♀, ABS: UCRCENT00305276]. Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 2.viii.2005, P. & M. Arnaud, malaise trap [1 ♀, ABS: UCRCENT00305280]. Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 21.ii.1979, H.V. Weems, Jr. & Sylvia Halkin, insect flight trap [2 ♀, FSCA: UCRCENT00241380– 81]. Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 24.x.1998, M. Deyrup, west side of LK. Annie, area of chironomid and spider abundance, yellow bowl trap [2 ♀, ABS: UCRCENT00305277–78]. Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 27.x.1998, M. Deyrup, west side of LK. Annie, area of chironomid and spider abundance, yellow bowl trap [1 ♀, ABS: UCRCENT00305279]. Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 9.vii.1979, H.V. Weems, Jr. & Cathy W. Harris, insect flight trap [1 ♀, FSCA: UCRCENT00241382]. Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 9.viii.2005, P. & M. Arnaud, malaise trap [1 ♀, ABS: UCRCENT00305271]. Archbold Bio. Research Station, 27°11'15"N, 81°20'26"W, 11.iii.2010, M. Deyrup, yellow sand scrub, site 6, flight trap [1 ♀, ABS: UCRCENT00305269]. Archbold Biol. Station, 27°10'54"N, 81°21'06"W, 1–18.ix.1987, D.B. Wahl [1 ♀, EMUS: UCRCENT00476388]. Archbold Biol. Station, 27°10'54"N, 81°21'06"W, 29.xii.1987 – 4.i.1988, D.B. Wahl [1 ♀, EMUS: UCRCENT00476389]. Archibold Biol. Stat., Lk. Placid, 27°10'54"N, 81°21'06"W, 4.iv.1986, M. Deyrup [1 ♀, ABS: UCRCENT00305349]. Archibold Biol. Stat., Lk. Placid, 27°10'54"N, 81°21'06"W, 9.iv.1986, M. Deyrup [1♂, ABS: UCRCENT00305355]. Lake Placid, Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 10.iii.1986, M. Deyrup, malaise trap [3 ♀, UCRC: UCRCENT00251456, UCRCENT00251460, UCRCENT00251462]. Lake Placid, Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 10.xi.1986, M. Deyrup, malaise trap [1 ♀, UCRC: UCRCENT00251455]. Lake Placid, Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 17.xi.1986, M. Deyrup, malaise trap [4 ♀, UCRC: UCRCENT00237634, UCRCENT00251452–53, UCRCENT00251458]. Lake Placid, Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 18–24.viii.1987, BRC Hym. team [1 ♀, UCRC: UCRCENT00251451]. Lake Placid, Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 22.xii.1992, M. Deyrup, trail 45, malaise trap [1 ♀, ABS: UCRCENT00305273]. Lake Placid, Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 5.xi.1986, M. Deyrup, malaise trap [1 ♀, UCRC: UCRCENT00251454]. Lake Placid, Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 6.ii.1985, M. Deyrup, malaise trap [1 ♀, UCRC: UCRCENT00251457]. Lake Placid, Archbold Bio. Research Station, 27°10'54"N, 81°21'06"W, 7.xi.1986, M. Deyrup, malaise trap [2 ♀, UCRC: UCRCENT00251459, UCRCENT00251461]. Indian River Co., 5 mi. S. Vero Beach, 27°43'07"N, 80°26'15"W, 1–13.ix.1983, J.H. Frank, woodland edge, malaise trap [1 ♀, FSCA: UCRCENT00241377]. 5 mi. S. Vero Beach, 27°43'07"N, 80°26'15"W, 13–23.ix.1983, J.H. Frank, woodland edge, malaise trap [1 ♀, FSCA: UCRCENT00241375]. Leon Co., Apalachicola National Forest, FS Rd 390: bottomland “mosquito hell”, 30°21'53"N, 84°40'46"W, 23–31.viii.2005, A. Vanderby, malaise trap [1 ♀, UCRC: UCRCENT00364793 (D3961)]. Hwy 375 4.0 km S. Bloxham, 28°44'29"N, 81°26'04"W, 4.vi.1987, J. Heraty, water oak, sweep [2 ♀, UCRC: UCRCENT00237632–33]. Hwy 375, 28°44'29"N, 81°26'04"W, 5.vi.1987, J. Heraty, on oak [1 ♀, UCRC: UCRCENT00237631]. Levy Co., 50 km. W. Archer, 28°34'10"N, 81°36'29"W, 20.vi–8.v.1986, LaSalle & Sharkey [1 ♀, CNC: UCRCENT00247584]. Manatee Co., Brandenton, 27°29'56"N, 82°34'29"W, 27.x–6.xii.1985, K. Patel & C. Yoshimoto, malaise trap/pan trap [1 ♀, CNC: UCRCENT00247578]. Marion Co., 8.0 km N Hwy 40 on 65 nr. Juniper Springs Rec. Area, 30m, 29°14'58"N, 81°43'38"W, 30.v.1987, J. Heraty, pine /palmetto forest [1 ♀, ABS: UCRCENT00305725]. Lake Eaton, 29°15'32"N, 81°52'03"W, 10.ix–2.x.1975, J. Wiley, malaise trap [1 ♀, CNC: UCRCENT00320860]. Monroe Co., 25°09'52"N, 81°00'02"W, 3.vi–27.viii.1986, S. & J. Peck, hammock, malaise trap & flight intercept trap [2 ♀, UCRC: UCRCENT00237635, UCRCENT00237639]. Bahia Honda Key, 24°39'52"N, 81°15'37"W, 25.x.1982, C. Potter & L. Stange [1 ♀, FSCA: UCRCENT00241376]. Orange Co., Apopka, Wekiwa Springs State Park. WSSPS-S- 1, 18m, 28°42'41"N, 81°27'45"W, 23–30.x.1993, Z. Prusak [3 ♀, UCFC: UCFC00165878–80]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 1–14.xi.1994, J.C. Longhurst, S.M. Fullerton, S Pine / Oak Scrub, MT [1 ♀, UCFC: UCFC00165853]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 10.x.1994, J C Longhurst, S.M. Fullerton, S Pine / Oak Scrub, MT [3 ♀, UCFC: UCFC00165840, UCFC00165845, UCFC00165872]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 12.x.1995, S.M. Fullerton, Summer Student, S Pine / Oak Scrub, Malaise Trap [1 ♀, UCFC: UCFC00123640]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 17.x.1994, J.C. Longhurst, S.M. Fullerton, S Pine / Oak Scrub, MT [1 ♀, UCFC: UCFC00165873]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 19.vii.1995, S.M. Fullerton, Summer Student, S Pine / Oak Scrub, Malaise Trap [2 ♀, UCFC: UCFC00165852, UCRC: UCRCENT00446708]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 26.vii.1995, S.M. Fullerton, Summer Students, S Pine / Oak Scrub, Malaise Trap [3 ♀, UCFC: UCFC00165846, UCFC00165851; UCRC: UCRCENT00446707]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 28.xi.1994, J.C. Longhurst and S.M. Fullerton, S Pine / Oak Scrub, Malaise Trap [1 ♀, UCFC: UCFC00123638]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 4.x.1994, J C Longhurst, S M Fullerton, S Pine / Oak Scrub, MT [1 ♀, UCFC: UCFC00165850]. Rock Springs Run State Reserve, 28°42'54"N, 81°27'00"W, 5.ix.1994, J.C. Longhurst, S.M. Fullerton, S Pine / Oak Scrub, MT [2 ♀, UCFC: UCFC00165868–69]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 1.viii.1997, S.M. Fullerton, Sand Pine- Rosemary Scrub, Pit Fall [2 ♀, UCFC: UCFC00126773–74]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 10.vii.1997, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [3 ♀, UCFC: UCFC00128693, UCFC00128695, UCFC00128697]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 10.vii.1997, S.M. Fullerton, Sand Pine, Rosemary Scrub, Pit Fall [1 ♀, UCFC: UCFC00126776]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 11.ix.1997, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [5 ♀, UCFC: UCFC00127403, UCFC00128740–43]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 11.ix.1997, S.M. Fullerton, Sand Pine, Rosemary Scrub, Pit Fall [1 ♀, UCFC: UCFC00126452]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 11.vi.1997, S.M. Fullerton, LLP-Sand Pine Turkey Oak, Malaise trap [1 ♀, UCFC: UCFC00126731]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 11.x.1999, T. Smith, P. Russell, S.M. Fullerton, Pond Pine Comm/ Dahoon Holly (S), MT [1 ♀, UCFC: UCFC00049770]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 12.vii.1996, S.M. Fullerton, LLP—Sand Pine, Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00124695]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 12.x.1995, S.M. Fullerton, LLP-S Pine / Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00165890]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 13.vi.1993, S.M. Fullerton, Cypress Dome, Malaise Trap [1 ♀, UCFC: UCFC00165884]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 14.vi.1996, S.M. Fullerton, LLP—Sand Pine, Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00124932]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 15.viii.1997, S.M. Fullerton, LLP—Sand Pine, Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00127327]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 16.x.1997, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [2 ♀, UCFC: UCFC00127590, UCFC00127613]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 17.ix.1997, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [3 ♀, UCFC: UCFC0 0 128802, UCFC00128811, UCFC00129183]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 17.vii.1997, S.M. Fullerton, LLP— Sand Pine, Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00127451]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 17.viii.1995, S.M. Fullerton, LLP—Sand Pine, Turkey Oak, Malaise Trap N [2 ♀, UCFC: UCFC00165886–87]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 18.ix.1997, S.M. Fullerton, Sand Pine-Rosemary Scrub, Pit Fall [3 ♀, UCFC: UCFC00126453–55]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 2.viii.1999, P. Russell and S. Fullerton, Pond Pine Comm., Dahoon Holly, Malaise Trap (S) [1 ♀, UCFC: UCFC00042608]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 2.x.1997, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [2 ♀, UCFC: UCFC00127694, UCFC00129191]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 20.vii.1995, S.M. Fullerton, LLP—Sand Pine, Turkey Oak, Malaise Trap N [1 ♀, UCFC: UCFC00165885]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 23.x.1997, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [2 ♀, UCFC: UCFC00128717, UCFC: UCFC00129163]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 24.vii.1997, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00128838]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 24.vii.1997, S.M. Fullerton, Sand Pine— Rosemary Scrub, Pit Fall [2 ♀, UCFC: UCFC00126771–72]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 25.ix.1997, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [4 ♀, UCFC: UCFC00128749–50, UCFC00129203, UCFC00129501]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 25.ix.1997, S.M. Fullerton, Sand Pine—Rosemary Scrub, Pit Fall [1 ♀, UCFC: UCFC00126456]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 25.vi.1997, S. M. Fullerton, LLP-Sand Pine / Turkey Oak, MT [1 ♀, UCFC: UCFC00129643]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 25.vi.1997, S.M. Fullerton, LLP-Sand Pine Turkey Oak, Malaise trap [1 ♀, UCFC: UCFC00129319]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 25.vi.1997, S.M. Fullerton, Long Leaf Pine—Saw Palmetto, Malaise trap [1 ♀, UCFC: UCFC00126699]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 25.xi.1997, S.M. Fullerton, Lang Leaf Pine / Saw Palmetto, MT [1 ♀, UCFC: UCFC00129499]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 27.ix.1996, S.M. Fullerton, LLP—Sand Pine, Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00125416]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 27.ix.1999, P. Russell and S. Fullerton, Pond Pine Comm., Dahoon Holly, Malaise Trap (S) [1 ♀, UCFC: UCFC00042799]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 27.viii.1997, S.M. Fullerton, LLP- Sand Pine, Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00127824]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 27.xi.1997, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [2 ♀, UCFC: UCFC001 28309, UCFC: UCFC00129215]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 30.v.1996, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [2 ♀, UCFC: UCFC00124603–04]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 30.v.1997, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00126615]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 30.x.1996, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00125263]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 30.x.1997, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00127345]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 31.v.1996, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00124644]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 4.ix.1997, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00128241]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 5.vii.1996, S.M. Fullerton, LLP-Sand Pine Turkey Oak, Malaise trap [1 ♀, UCFC: UCFC00124801]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 5.x.1995, S.M. Fullerton, LLP—Sand Pine, Turkey Oak, Malaise Trap [2 ♀, UCFC: UCFC00123314, UCFC00165889]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 5.x.1999, P. Russell and S. Fullerton, Pond Pine Comm., Dahoon Holly, Malaise Trap (S) [1 ♀, UCFC: UCFC00042762]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 6.ix.1996, S.M. Fullerton, LLP—Sand Pine, Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00125006]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 7.ix.1995, S.M. Fullerton, LLP—Sand Pine, Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00124306]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 7.ix.1995, S.M. Fullerton, LLP—Sand Pine, Turkey Oak, Malaise Trap N [1 ♀, UCFC: UCFC00165888]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 7.viii.1997, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00128680]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 8.xi.1996, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [5 ♀, UCFC: UCFC0 0 125274, UCFC00125280, UCFC00125284–85, UCFC: UCFC00125304]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 9.x.1997, S. M. Fullerton, LLP-Sand Pine / Turkey Oak, MT [1 ♀, UCFC: UCFC00129198]. UCF, Orlando, 28°36'07"N, 81°12'00"W, 9.x.1997, S.M. Fullerton, LLP-Sand Pine, Turkey Oak, Malaise Trap [2 ♀, UCFC: UCFC00129196, UCFC00129202]. UCF, Orlando, MacKay Tract, 28°35'27"N, 81°13'01"W, 17.v.1999, P. Russell and S. Fullerton, Sawgrass Marsh, Red Maple, Malaise Trap [1 ♀, UCFC: UCFC00039458]. UCF, Orlando, MacKay Tract, 28°35'27"N, 81°13'01"W, 17.viii.1999, P. Russell and S. Fullerton, Sawgrass Marsh, Red Maple, Malaise Trap [2 ♀, UCFC: UCFC00047399, UCFC00047534]. UCF, Orlando, MacKay Tract, 28°35'27"N, 81°13'01"W, 2.vi.1999, P. Russell and S. Fullerton, Sawgrass Marsh, Red Maple, Malaise Trap [1 ♀, UCFC: UCFC00037649]. UCF, Orlando, MacKay Tract, 28°35'27"N, 81°13'01"W, 24.v.1999, P. Russell and S. Fullerton, Sawgrass Marsh, Red Maple, Malaise Trap [1 ♀, UCFC: UCFC00040140]. Wekiwa Sp. St. Pk., Burn Zone WS-14a, 28°44'32"N, 81°30'33"W, 5.viii.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, at ground level [1 ♀, UCFC: UCFC00141265]. Polk Co., Lake Wales Ridge State For., 27°39'40"N, 81°23'47"W, 6.v.2009, H. Otte & M. Deyrup, fl. scrub, site 3., flight trap w. pan [1 ♀, ABS: UCRCENT00305274]. Sarasota Co., Fruitville, 9m, 27°19'47"N, 82°27'27"W, 11.viii.1930, R.H. Beamer [1 ♀, SEMC: UCRCENT00404264]. MCC-Venice Campus, 3m, 27°02'30"N, 82°19'04"W, 20.i.1997, K.J. Maharay and S.M. Fullerton, Long Leaf Pine—Saw Palmetto, Malaise Trap [1 ♀, UCFC: UCFC00125581]. MCC-Venice Campus, 3m, 27°02'30"N, 82°19'04"W, 27.i.1997, K.J. Maharay and S.M. Fullerton, Long Leaf Pine—Saw Palmetto, Malaise Trap [1 ♀, UCFC: UCFC00125580]. Seminole Co., Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 1.iv.2000, Trevor Smith, Scrub Oak / Saw Palmetto (burned), Malaise Trap [3 ♀, UCFC: UCFC00053489, UCFC00053492, UCFC00053850]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 1.x.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (unburned), MT [1 ♀, UCFC: UCFC00078755]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 15.jul.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (Burned), MT [1 ♀, UCFC: UCFC00072228]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 15.vi.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (Burned), MT [1 ♀, UCFC: UCFC00072196]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 15.vii.2000, T. Smith, P. Russell and S.M. Fullerton, Scrub Oak / Saw Palmetto (burned), Malaise Trap [1 ♀, UCFC: UCFC00072265]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 16.xii.2000, T. Smith, P. Russell and S.M. Fullerton, Scrub Oak / Saw Palmetto (unburned), Malaise Trap [1 ♀, UCFC: UCFC00085333]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 17.iv.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (burned), MT [1 ♀, UCFC: UCFC00060318]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 17.ix.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (Burned), MT [1 ♀, UCFC: UCFC00075625]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 17.ix.2000, T. Smith, P. Russell, S.M. Fullerton, Scrub Oak / Saw Palmetto (burned), Malaise Trap [1 ♀, UCFC: UCFC00071800]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 18.xi.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (unburned), MT [1 ♀, UCFC: UCFC00080546]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 19.ii.2000, T. Smith, Scrub Oak / Saw Palmetto (Unburned), MT [1 ♀, UCFC: UCFC00053661]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 2.ix.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (Burned), MT [1 ♀, UCFC: UCFC00070940]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 20.v.2000, T. Smith, P. Russell and S.M. Fullerton, Scrub Oak / Saw Palmetto (unburned), Malaise Trap [1 ♀, UCFC: UCFC00064232]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 20.v.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (unburned), MT [1 ♀, UCFC: UCFC00064095]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 20.v.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (burned), MT [1 ♀, UCFC: UCFC00061568]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 20.viii.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (Unburned), MT [1 ♀, UCFC: UCFC00074213]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 22.vii.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (Unburned), MT [1 ♀, UCFC: UCFC00072473]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 22.vii.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (burned), MT [1 ♀, UCFC: UCFC00072564]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 22.x.2000, T. Smith, Scrub Oak / Saw Palmetto (Unburned), MT [1 ♀, UCFC: UCFC00080633]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 24.ix.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (burned), MT [1 ♀, UCFC: UCFC00076087]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 27.v.2000, T. Smith, P. Russell and S.M. Fullerton, Scrub Oak / Saw Palmetto (burned), Malaise Trap [1 ♀, UCFC: UCFC00064465]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 29.iv.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (Burned), MT [3 ♀, UCFC: UCFC00059969, UCFC00060005, UCFC00060009]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 3.xii.2000, T. Smith, P. Russell and S.M. Fullerton, Scrub Oak / Saw Palmetto (unburned), Malaise Trap [1 ♀, UCFC: UCFC00081448]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 30.vii.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (unburned), MT [1 ♀, UCFC: UCFC00072839]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 6.v.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (Burned), MT [1 ♀, UCFC: UCFC00061092]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 8.vii.2000, T. Smith, P. Russell, S.M. Fullerton, Scrub Oak / Saw Palmetto (Burned), MT [1 ♀, UCFC: UCFC00068285]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 8.x.2000, T. Smith, P. Russell and S.M. Fullerton, Scrub Oak / Saw Palmetto (unburned), Malaise Trap [1 ♀, UCFC: UCFC00078276]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 8.x.2000, T. Smith, P. Russell and S.M. Fullerton, Scrub Oak / Saw Palmetto (burned), Malaise Trap [1 ♀, UCFC: UCFC00078500]. Econ Wild. Area, 12m, 28°36'49"N, 81°10'04"W, 8.x.2000, T. Smith, P. Russell, S.M Fullerton, Scrub Oak / Saw Palmetto (unburned), MT [1 ♀, UCFC: UCFC00078311]. Econ Wilderness Area, 12m, 28°36'49"N, 81°10'04"W, 19.ii.2000, Trevor Smith, Scrub Oak / Saw Palmetto (burned), Malaise Trap [2 ♀, UCFC: UCFC00047278, UCFC00047285]. Econ Wilderness Area, 12m, 28°36'49"N, 81°10'04"W, 19.ii.2000, Trevor Smith, Scrub Oak / Saw Palmetto (unburned), Malaise Trap [1 ♀, UCFC: UCFC00053610]. Econ Wilderness Area, 12m, 28°36'49"N, 81°10'04"W, 25.iii.2000, Trevor Smith, Scrub Oak / Saw Palmetto (burned), Malaise Trap [1 ♀, UCFC: UCFC00054292]. Econ Wilderness Area, 12m, 28°36'49"N, 81°10'04"W, 8.iv.2000, Trevor Smith, Scrub Oak / Saw Palmetto (burned), Malaise Trap [1 ♀, UCFC: UCFC00054734]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 14.iv.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Trap, at ground level [1 ♀, UCFC: UCFC00133753]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 14.v.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, 3 ft above ground [1 ♀, UCFC: UCFC00138453]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 14.v.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, 1 ft above ground [1 ♀, UCFC: UCFC00138271]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 14.x.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, at ground level [3 ♀, UCFC: UCFC00142081, UCFC00142968, UCFC00142972]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 16.vii.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, 1 ft above ground [2 ♀, UCFC: UCFC00144764, UCFC00144780]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 16.xii.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, 3 ft above ground [3 ♀, UCFC: UCFC00144632, UCFC00144646, UCFC00144651]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 16.xii.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, 1 ft above ground [1 ♀, UCFC: UCFC00144774]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 16.xii.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, at ground level [1 ♀, UCFC: UCFC00144688]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 16.xii.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, 3 ft above ground [1 ♀, UCFC: UCFC00144845]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 18.xi.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, 1 ft above ground [2 ♀, UCFC: UCFC00144310, UCFC00144311]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 18.xi.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, at ground level [1 ♀, UCFC: UCFC00144331]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 19.viii.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, at ground level [1 ♀, UCFC: UCFC00141858]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 2.ix.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, at ground level [1 ♀, UCFC: UCFC00142133]. Lower Wekiva River St. Pres., Burn Zone LW- 10, 28°49'28"N, 81°24'26"W, 22.vii.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, 3 ft above ground [1 ♀, UCFC: UCFC00140573]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 27.v.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, at ground level [1 ♀, UCFC: UCFC00134723]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 27.v.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, at ground level [1 ♀, UCFC: UCFC00134656]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 29.iv.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, 3 ft above ground [1 ♀, UCFC: UCFC00136118]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 29.iv.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, at ground level [3 ♀, UCFC: UCFC00136011, UCFC: UCFC00136018–19]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 29.iv.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, 3 ft above ground [1 ♀, UCFC: UCFC00136110]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 30.xii.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, at ground level [1 ♀, UCFC: UCFC00144926]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 30.xii.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, at ground level [2 ♀, UCFC: UCFC00144944, UCFC00144950]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 4.xi.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, at ground level [1 ♀, UCFC: UCFC00143696]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 4.xi.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, 1 ft above ground [3 ♀, UCFC: UCFC00144025, UCFC00144029, UCFC00144031]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 4.xi.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, 1 ft above ground [3 ♀, UCFC: UCFC00143991, UCFC00144004, UCFC00144018]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 4.xi.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, 3 ft above ground [1 ♀, UCFC: UCFC00143782]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 4.xi.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, at ground level [4 ♀, UCFC: UCFC00143724, UCFC00143846, UCFC: UCFC00143880, UCFC00143887]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 4.xi.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, at ground level [4 ♀, UCFC: UCFC00143724, UCFC00143846, UCFC00143880, UCFC00143887]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 8.vii.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, at ground level [1 ♀, UCFC: UCFC00140054]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 8.vii.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, 3 ft above ground [1 ♀, UCFC: UCFC00139874]. Lower Wekiva River St. Pres., Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 8.vii.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, 1 ft above ground [1 ♀, UCFC: UCFC00140156]. Lower Wekiva River St. Pres., Burn Zone LW-10, S39 T19S R29E, 11m, 28°49'30"N, 81°24'27"W, 18.xi.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, 3 ft above ground [1 ♀, UCFC: UCFC00144150]. Lower Wekiva River St. Pres., Burn Zone LW-10, S39 T19S R29E, 11m, 28°49'30"N, 81°24'27"W, 18.xi.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, at ground level [5 ♀, UCFC: UCFC00144245, UCFC00144247, UCFC00144253, UCFC00144256, UCFC00144262]. Lower Wekiva River St. Pres., Burn Zone LW-10, S39 T19S R29E, 11m, 28°49'30"N, 81°24'27"W, 3.iv.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Flor. Yel. Pan Trap, 3 ft above ground [1 ♀, UCFC: UCFC00137801]. Lower Wekiva River St. Pres., Burn Zone LW-10, S39 T19S R29E, 11m, 28°49'30"N, 81°24'27"W, 3.iv.2001, P. Russell and S. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, at ground level [1 ♀, UCFC: UCFC00137622]. Lower Wekiva River St. Preserve Burn Zone, 28°49'48"N, 81°24'36"W, 30.ix.2001, P.J Russell, S.M. Fullerton, Longleaf Pine-Turkey Oak, Malaise [1 ♀, UCFC: UCFC00114198]. Lower Wekiva River St. Preserve, Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 2.xii.2001, P.J. Russell and S.M. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, at ground level [2 ♀, UCFC: UCFC00144454, UCFC00144508]. Lower Wekiva River St. Preserve, Burn Zone LW-10, 28°49'28"N, 81°24'26"W, 2.xii.2001, P.J. Russell and S.M. Fullerton, LLP-Turkey Oak, Yellow Pan Trap, 1 ft above ground [1 ♀, UCFC: UCFC00144600]. Lower Wekiva River St. Preserve, Burn Zone LW-10, S39 T19S R29E, 11m, 28°49'30"N, 81°24'27"W, 28.x.2001, P.J. Russell and S.M. Fullerton, LLP- Turkey Oak, Yellow Pan Trap, at ground level [1 ♀, UCFC00143523]. Lower Wekiva River St. Preserve, Burn Zone LW-5, 28°49'28"N, 81°24'26", 16.ix.2001, P.J. Russell and S.M. Fullerton, Longleaf Pine-Turkey Oak, Malaise Trap [2 ♀, UCFC: UCFC00111952, UCFC00112037]. Lower Wekiva River St. Preserve, Burn Zone LW- 5, 28°49'28"N, 81°24'26"W, 13.iv.2001, P.J. Russell and S.M. Fullerton, Longleaf Pine—Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00109390]. Lower Wekiva River St. Preserve, Burn Zone LW-5, 28°49'28"N, 81°24'26"W, 13.v.2001, P.J. Russell and S.M. Fullerton, Longleaf Pine—Turkey Oak, Malaise Trap [2 ♀, UCFC: UCFC00109905, UCFC00109974]. Lower Wekiva River St. Preserve, Burn Zone LW-5, 28°49'28"N, 81°24'26"W, 18.xi.2001, P.J. Russell and S.M. Fullerton, Longleaf Pine—Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00116462]. Lower Wekiva River St. Preserve, Burn Zone LW-5, 28°49'28"N, 81°24'26"W, 2.xii.2001, P.J. Russell and S.M. Fullerton, Longleaf Pine—Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00118742]. Lower Wekiva River St. Preserve, Burn Zone LW-5, 28°49'28"N, 81°24'26"W, 21.i.2001, P.J. Russell and S.M. Fullerton, Longleaf Pine—Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00111427]. Lower Wekiva River St. Preserve, Burn Zone LW-5, 28°49'28"N, 81°24'26"W, 23.vi.2001, P.J. Russell and S.M. Fullerton, Longleaf Pine—Turkey Oak, Malaise Trap [1 ♀, UCFC: UCFC00110444]. Lower Wekiva River St. Preserve, Burn Zone LW-5, 28°49'28"N, 81°24'26"W, 26.v.2001, P.J. Russell and S.M. Fullerton, Longleaf Pine—Turkey Oak, Malaise Trap [2 ♀, UCFC: UCFC00110191, UCFC00110259]. Lower Wekiva River St. Preserve, Burn Zone LW-5, 28°49'28"N, 81°24'26"W, 28.iv.2001, P.J. Russell and S.M. Fullerton, Longleaf Pine—Turkey Oak, Malaise Trap [3 ♀, UCFC: UCFC00109754, UCFC00109774, UCFC00109786]. Oviedo, 16m, 28°40'21"N, 81°12'25"W, 10.x.1994, S.M. Fullerton, Rural Yard, Malaise [1 ♀, UCFC: UCFC00165858]. Oviedo, 16m, 28°40'21"N, 81°12'25"W, 5.x.1994, S.M. Fullerton, Rural Yard, Malaise [1 ♀, UCFC: UCFC00165859]. Taylor Co., Blue Springs Lake, 28°52'08"N, 81°23'37"W, 4.vi.1972, C.L. Smith, black light [1 ♀, UCRC: UCRCENT00237629]. Volusia Co., Holly Hill, 3m, 29°14'37"N, 81°02'15"W, 1.vi.1978, G.J. Umphrey [1 ♀, DEBU: UCRCENT00251198]. Walton Co., DeFuniak Springs, 30°43'16"N, 86°06'55"W, 27.x-17.xi.1991, Deyrup & Winegarner [1 ♀, ABS: UCRCENT00305272]. Georgia: Okeefenokee swamp, 32°35'49"N, 82°57'28"W, 26.viii.1939, D.E. Hardy [1 ♀, CNC: UCRCENT00252359]. Okeefenokee swamp, 32°35'49"N, 82°57'28"W, 31.viii.1939, D.E. Hardy [1 ♀, UCRC: UCRCENT00252354]. Charlton Co., Okeefenokee Nat. Wildlife Refuge, 32°35'49"N, 82°57'28"W, 8.ix.1976, C.L. Smith [1 ♀, UCRC: UCRCENT00252353]. McIntosh Co., Sapelo Island, 31°23'51"N, 81°16'43"W, 20.vi– 18.vii.1987, BRC Hym. team, open sand dunes, malaise trap [1 ♀, CNC: UCRCENT00247624]. Sapelo Island, 31°23'51"N, 81°16'43"W, 20.vii.1987, BRC Hym. team, live oak forest, FIT [1 ♀, UCRC: UCRCENT00252348]. Sapelo Island, 31°23'51"N, 81°16'43"W, 9.i–20.viii.1987, BRC Hym. team, shrub sand dunes, pan trap [1 ♀, UCRC: UCRCENT00252371]. Sapelo Island, 31°23'51"N, 81°16'43"W, 9.v–20.vi.1987, BRC Hym. team, open sand dunes, pan trap [4 ♀, CNC: UCRCENT00247620–23]. Ware Co., Okeefenokee Swamp, 30°37'07"N, 82°19'15"W, 27.vii.1939, A.T. Hardy [1 ♀, SEMC: UCRCENT00404265]. Okeefenokee Swamp, 30°37'07"N, 82°19'15"W, 27.vii.1939, R.H. Beamer [1 ♀, SEMC: UCRCENT00404266]. Louisiana: Calcasieu, Lake Charles, 30°13'36"N, 93°13'03"W, 12.iv.1985, D. Wheeler, Pheidole in wood [4 ♀, UCRC: UCRCENT00252355–58]. North Carolina: Hatteras Is. Buxton Forest, 35°15'33"N, 75°31'35"W, 16.viii.1980, L. Masner [4 ♀, CNC: UCRCENT00247616–19]. Carteret Co., 2006 Emeline Pl., Moorehead City, 34°43'48"N, 76°43'12"W, 20– 27.vii.2000, Mark Wuenschel [3 ♀, UCRC: UCRCENT00414212–14]. 2006 Emiline Place, Moorehead City, 34°43'49"N, 76°43'48"W, 20–27.viii.2000, Mark Wuenschel [2 ♀, UCRC: UCRCENT00091467, UCRCENT00091468 (D1005)]. Texas: Bandera Co., Lost Maples St. Pk., 29°49'12"N, 99°34'12"W, 18.viii.1988, G. Zolnerowich [2 ♀, TAMU: UCRCENT00243220–21]. Montgomery Co., Jones State Forest 8 mi. S Conroe, 30°12'54"N, 95°22'12"W, 18–24.v.1987, Wharton Wang. Praetorius [2 ♀, TAMU: UCRCENT00243385–86]. Jones State Forest 8 mi. S Conroe, 30°12'54"N, 95°22'12"W, 25–31.v.1987, Wharton Wang. Praetorius [3 ♀, TAMU: UCRCENT00426501–03]. Jones State Forest, 8 mi. S Conroe, 57m, 30°13'20"N, 95°29'20"W, 1– 7.vi.1987, Wharton, Wang, and Praetorius [1 ♀, TAMU: UCRCENT00243223]. Jones State Forest, 8 mi. S Conroe, 57m, 30°13'20"N, 95°29'20"W, 1–7.vii.1987, Wharton, Wang, and Praetorius [3 ♀, TAMU: UCRCENT00426504–06]. Jones State Forest, 8 mi. S Conroe, 57m, 30°13'20"N, 95°29'20"W, 1.vii–6.ix.1987, Wharton, Steck. and Carroll [1 ♀, TAMU: UCRCENT00426507]. Nacogdoches Co., Steven F. Austin Experimental Forest, Angelina National Forest, 12 km N Nacogdoches, 31°42'00"N, 94°40'00"W, 16.v– 15.vi.2003, N. Schiff & C. Rudolph [2 ♀, TAMU: UCRCENT00479288, UCRC: UCRCENT00446773]. Tyler Co., 3 miles S. Warren, 32°21'18"N, 97°25'07"W, 22.v.1984, J. B. Woolley [1♂, TAMU: UCRCENT00426510]. Walker Co., Huntsville St. Pk., 30°52'23"N, 95°29'45"W, 15.viii.1986, J. Heraty, open oak forest [1 ♀, UCRC: UCRCENT00252249]. Huntsville St. Pk., 30°52'23"N, 95°29'45"W, 17.viii.1986, J. Heraty, Oak forest [1 ♀, UCRC: UCRCENT00252240]. Stubblefield Lake nature tr., 30°33'56"N, 95°38'50"W, 19.v.1999, J.M. Heraty, beech / pine [1♂ 8 ♀, UCRC: UCRCENT00091378–81, UCRCENT00091383–86, UCRC: UCRCENT00092177]. Stubblefield Lake nature tr., 30°33'56"N, 95°38'50"W, 4.x.1986, J.M. Heraty, ex: Pheidole dentata [2 ♀, UCRC: UCRCENT00408457 (D3764), UCRENT00408458 (D3765)]. Stubblefield Lk Tr., 29°01'49"N, 95°19'49"W, 4.x.1986, J. Heraty [23 ♀, UCRC: UCRCENT00237485, UCRCENT00237637–38, UCRCENT00251464, UCRCENT00251486–87, UCRCENT00252216–19, UCRCENT00252221–28, UCRCENT00252230–32, UCRCENT00252243, UCRCENT00278550]. Stubblefield Lk Tr., 30°33'45"N, 95°39'07"W, 4.x.1986, J. Heraty, in log [3 ♀, UCRC: UCRCENT00252238, UCRCENT00414245, UCRCENT00414248]. Walker Co., Stubblefield Lk Tr., 29°01'49"N, 95°19'49"W, 4.x.1986, J. Heraty [1 ♀, UCRC: UCRCENT00252244]. Virginia: Essex Co., 1mi. S.E. Dunnsville, 37°51'12"N, 76°46'19"W, 10–26.viii.1991, D.R. Smith, malaise trap [2 ♀, USNM: UCRCENT00247797–98]. Virginia Beach, 36°50'23"N, 76°02'44"W, E.S.G. Titus [1 ♀, USNM: UCRCENT00248654].

Planidia. USA: Texas: Walker Co., Huntsville St. Pk. , 30°52'23"N, 95°29'45"W, 14.viii.1986, J. Heraty, open oak forest, host: on leaves of water oak [7 slides, UCRC: UCRCENT00478805–11 ] . Stubblefield Lk Trail , 30°52'23"N, 95°29'45"W, 4.x.1986, J. Heraty, open oak forest, nest of Pheidole dentata under broken log [2 slides, UCRC: UCRCENT00478812–13 ] . Orasema evansi ? (planidia): Costa Rica: Puntarenas: Coto , 8m, 8°32'08"N, 83°02'05"W, 29.x.1956, C.E. Palm, in citrus leaf, host plant: citrus [ USNM: UCRCENT00416825 ] . Coto , 8m, 8°32'08"N, 83°02'05"W, 5.iii.1956, R.V. Roig, on mango leaves [ USNM: UCRCENT00416822–24 , UCRCENT00425532–35 ] .

Pupae. USA: Florida: Holly Hill, 29°14'37"N, 81°02'15"W, 1.vi.1978, G.J. Umphrey [2 ♀, UCRC: UCRCENT00237565–66]. Alachua Co., Gainesville, 29°39'06"N, 82°19'29"W, 13.vi.1996, L. Davis, H96-D34, host: Pheidole dentata [4 ♀ 1♂, FSCA: UCRCENT00411923–27]. Hague dairy forest, 29°47'28"N, 82°25'07"W, 10.ix.2011, John Sivinski, in rotted pine branch, host: Pheidole dentata [1 ♀, EMUS: UCRCENT00476412]. Putnam Co., Ordway Swisher Biol. Sta., 29°42'08"N, 81°59'29"W, 23.vi.2013, John Sivinski, host: Pheidole dentata [1 ♀, EMUS: UCRCENT00476399].

Etymology. The specific epithet is in honor of Howard Ensign Evans, who discussed this species in his book “Life on a Little Known Planet” ( Evans 1968: 245).

Figures

FIGURE 3. Maximum likelihood phylogeny of the Orasema stramineipes species group. Branch color indicates major ecological regions, separating Neotropical (green), Nearctic (red) and southern subtropical to temperate (blue). Numbers associated with branchesindicate bootstrap valuesabove 50.Blue lettersrefer to host associationswithin a clade:P = Pheidole, S = Solenopsis, W = Wasmannia. Species names are associated with the molecular voucher number and sex; vertical tan lines group specimenscollected from the same locality.Inset figures are of the dorsalmesosoma and species distribution maps based on all material examined.

FIGURES 1, 2. Orasema stramineipes group biology. 1. Orasema freychei (Argentina) ovipositing on underside of leaf of Tecoma stans; leaf covered with oviposition scars. 2. Orasema evansi (Costa Rica) oviposition scars on banana (USDA-SEL archive photo).

FIGURES 61–66. Orasema evansi. ♀: form from banana in Costa Rica: 61. Habitus. 62. Head. 63. Antenna, with F2 and F6– F7 inset.64. Mesosoma,dorsal view.65. Axillula. 66. Stigma and postmarginal vein.

FIGURES 67–68. Orasema evansi. ♀: form from banana in Costa Rica: 67. Propodeum.68. Petiole.

FIGURES 69–74. Orasema evansi. ♀: USA form: 69. Habitus. 70. Head. 71. Antenna, with F2 and F6–F7 inset. 72. Mesosoma, dorsal view. 73. Axillula. 74. Stigma andpostmarginal vein.

FIGURES 75–76. Orasema evansi. ♀: USA form: 75.Propodeum.76. Petiole.

FIGURE4. Orasema evansi(Texas), immature stages and hosts:A.Stalkedegg.B.Unfed planidium(first instar),D. Pheidole dentata larva with internal and swollen (fed) planidium indicated by arrow; posterior apex open to outside by entrance hole in cuticle.E. P.dentata pupa with external planidium(larval exuvium shed).F. O. evansi2nd instar larva.G. P.dentata pupa with third instar O. evansi larva attached. H. O. evansi prepupa. I. O. evansi pupa. K. Metasoma of pupa Orasema minutissima (Puerto Rico):C. Unfed planidium larvae.J. Unparasitized, but fed-upon, pupa of P.dentata (psithergate).

FIGURES 17–18. Orasema arimbome. ♀: 17. Propodeum, cal = propodeal callus, mtn = metanotum, ppd = propodeum. 18. Petiole (pet), ptc = anterior petiolar carina.